Determinants of urban tree canopy in residential neighborhoods: Household characteristics, urban form, and the geophysical landscape

The aesthetic, economic, and environmental benefits of urban trees are well recognized. Previous research has focused on understanding how a variety of social and environmental factors are related to urban vegetation. The aim is often to provide planners with information that will improve residential neighborhood design, or guide tree planting campaigns encouraging the cultivation of urban trees. In this paper we examine a broad range of factors we hypothesize are correlated to urban tree canopy heterogeneity in Salt Lake County, Utah. We use a multi-model inference approach to evaluate the relative contribution of these factors to observed heterogeneity in urban tree canopy cover, and discuss the implications of our analysis. An important contribution of this work is an explicit attempt to account for the confounding effect of neighborhood age in understanding the relationship between human and environmental factors, and urban tree canopy. We use regression analysis with interaction terms to assess the effects of 15 human and environmental variables on tree canopy abundance while holding neighborhood age constant. We demonstrate that neighborhood age is an influential covariate that affects how the human and environmental factors relate to the abundance of neighborhood tree canopy. For example, we demonstrate that in new neighborhoods a positive relationship exists between street density and residential tree canopy, but the relationship diminishes as the neighborhood ages. We conclude that to better understand the determinants of urban tree canopy in residential areas it is important to consider both human and environmental factors while accounting for neighborhood age.     

Tree diversity,distribution, history and change in urban parks: studies in Bangalore,India

Urban parks constitute critical biodiversity hotspots in crowded, concrete-dominated city environments. Despite the importance, they remain little researched. This paper assesses the biodiversity and distribution of trees in urban parks in the southern Indian city of Bangalore. 127 plots were used to survey tree distribution in parks across the city. The distribution is largely dominated by a few common species. The proportion of exotic species was very high, with 77% of trees belonging to introduced species. Park history had an impact on distribution. Old parks had fewer but larger trees, and greater species diversity compared to recently established parks. Old parks also differed in species composition, having a greater proportion of large canopy trees compared to young parks. Examination of size distributions revealed that large canopied species were gradually being phased out, and replaced by narrow and medium sized tree species which are easier to maintain, but which may not provide the same environmental and ecological benefits. Greater attention requires to be paid to the selection of trees in cities, not just with a view to easy maintenance as is currently the case, but to select an appropriate mix of trees that supports biodiversity and maximizes environmental and ecosystem services.     

The impact of urban forest structure and its spatial location on urban cool island intensity

Urban forest can help decrease land surface temperature (LST) and create urban cooling effect (UCI) to mitigate urban heat island (UHI). However, it is still unclear how urban forest structure and its location affect UCI, particularly under different seasons. In this study, with plot-based urban forest structure and UCI intensity extracted from Landsat-7 Enhanced Thematic Mapper Plus (ETM+) thermal data, we first conducted correlation analyses between UCI and different forest structures (crown closure, tree height, leaf area index, basal area, stem density and diameter at breast height, etc.) and spatial location (distances from buildings and from water bodies and elevation) attributes, and we then carried out quantitative regression analyses between them. Our results indicate that (1) Urban forest could create “urban cool islands”, which were higher in summer than those in autumn.(2) UCI could be significantly affected by urban forest structural attributes, especially by crown closure and LAI. All urban forest structural attributes had positive linear relationships with UCI except for LAI and basal area which had positive non-linear relationships with UCI.; (3) UCI in urban forest could also be affected by its spatial location but not by its elevation. The UCI non-linearly decreased with decreasing distance from buildings and with increasing distance from water bodies. The threshold values of DB for significantly affecting UCI variation is approximately between 100 m and 300 m in summer and autumn, respectively; and (4) the relationships between UCI and urban forest structure and its location attributes were complex and seasonal dependent. Urban forest attributes had greater effects on increasing UCI in summer than those in autumn. These findings would deepen our understanding of interactions between UCI and urban forest attributes and provide urban planners with useful information about how to design urban forest to effectively mitigate UHI effects.     

Modelling changes in dimensions,health status,and arboricultural implications for urban trees

Canberra, the capital of Australia since 1911, has been developed into a modern city from its original site on a nearly treeless plain. Today the city has about 300,000 inhabitants and 500,000 trees. The authors were requested by the managers of the urban public tree resource to survey their asset and to develop a computer-based system that would aid them in anticipating future maintenance requirements and its costs. This paper reports on our response. We have surveyed 3,000 streets and parks in the city, noting the species, number, and condition of every public space tree. We have also obtained the dimensions of sample trees, noting their total height, maximum crown width, height of maximum crown width, diameter at maximum crown width, and height at crown break. A management system has been developed using Microsoft Access^TM.Using standard regression techniques available on the package JMP^TM, we found that total tree height was related to age for all species and that all other parameters of interest were related to height or transformed values of height. We assumed a sigmoidal growth curve and calibrated 114 height/age curves to cover the 165,000 trees of the 340 species we have in our database. As well we used the data on tree condition to determine the rate at which populations change from healthy to stressed.By interviewing foremen and supervisors we were able to determine the maintenance treatments carried out in Canberra, the equipment used, and the number of trees that can be treated in a day, for each type of operation.The management system can be used to display the current inventory for each street or park, by suburb, in the database. It can also be used to model future increases in size or crown condition, to predict the operations that will be required as a consequence of tree growth or crown deterioration, and finally, by applying multipliers to equipment and personnel, to estimate the future costs of tree maintenance. Managers can use the system to anticipate problems such as uneven expenditure requirements in future years.     

Improving confidence in tree species selection for challenging urban sites: a role for leaf turgor loss

High species diversity is argued to be the most important requisite for a resilient urban forest. In spite of this, there are many cities in the northern hemisphere that have very limited species diversity within their tree population. Consequently, there is an immense risk to urban canopy cover, if these over-used species succumb to serious pests or pathogens. Recognition of this should motivate the use of less commonly used species. Analysis of plant traits, such as the leaf water potential at turgor loss (Ψ_P0), can provide useful insights into a species’ capacity to grow in warm and dry urban environments. Therefore, the aim of this study was to evaluate Ψ_P0 of 45 tree species, the majority of which are rare in urban environments. To help evaluate the potential for using Ψ_P0 data to support future decision-making, a survey of professionals engaged with establishing trees in urban environments was also used to assess the relationship between the measured Ψ_P0 and the perceived drought tolerance of selected species. This study demonstrates that Ψ_P0 gives strong evidence for a species’ capacity to tolerate dry growing conditions and is a trait that varies substantially across species. Furthermore, Ψ_P0 was shown to closely relate to the experience of professionals involved in establishing trees in urban environments, thus providing evidence of its practical significance. Use of plant traits, such as Ψ_P0, should, therefore, give those specifying trees confidence to recommend non-traditional species for challenging urban environments.     

Habitat richness and tree species richness of roundabouts: effects on site selection and the prevalence of arboreal caterpillars

This study examined the relationship between the habitat and tree species richness of roundabouts and the abundance and species richness of tree-dwelling caterpillars, and thus the potential of urban roundabouts to support breeding populations. Total tree species number on sites was related to an increase in caterpillar abundance and a greater number of habitats was associated with a higher number of caterpillar species. An increase in the total number of trees was not correlated with a greater abundance of tree-feeding Lepidoptera; nor were native trees found to support proportionally more caterpillar species than all tree species, including introduced. This could be due to differences in species accumulation, the prevalence of generalist species or a statistical sampling effect. The occupancy and average abundance of caterpillar species was generally low, which does not support the theory that Lepidoptera in urban areas tend towards mono-dominance. It may, however, represent the presence of ‘accidental’ species on roundabouts. Habitat management and planting to increase tree species diversity and density on roundabouts has the potential to increase the species diversity and abundance of arboreal Lepidoptera, and increase abundance of butterfly and moth urban populations.     

Nesting ecology of mourning doves in an urban landscape

The expansion of urban areas into native habitat can have profound effects on avian populations and communities, yet little is known regarding the effects of urban features on avian reproductive success. The objective of this study was to examine the reproduction of an urban-enhanced species, the mourning dove, to determine how tree and urban landscape features affect nest-site selection and nest success. Mourning dove nests were located by systematically searching potential nest sites on a weekly basis from late-March through mid-September in 2003 and 2004. A total of 1,288 mourning dove nests were located and monitored on the Texas A&M University Campus. Of these nests, 337 (26.6%) were successful (fledged, ≥1). An equal number of potential nest sites were randomly generated in ArcGIS and assigned to non-nest trees to evaluate habitat variables associated with nest-site selection. Mourning dove nests were located in trees with a larger canopy diameter and diameter at breast height (DBH) than the computer generated potential nests and nest trees were located closer to roads and farther from buildings than non-nest trees. Within the study area, nest success was predominately influenced by the proximity of urban features with successful nests being located closer to roads and farther from buildings than unsuccessful nests.     

Tree health mapping with multispectral remote sensing data at UC Davis, California

Tree health is a critical parameter for evaluating urban ecosystem health and sustainability. Traditionally, this parameter has been derived from field surveys. We used multispectral remote sensing data and GIS techniques to determine tree health at the University of California, Davis. The study area (363 ha) contained 8,962 trees of 215 species. Tree health conditions were mapped for each physiognomic type at two scales: pixel and whole tree. At the pixel scale, each tree pixel within the tree crown was classified as either healthy or unhealthy based on vegetation index values. At the whole tree scale, raster based statistical analysis was used to calculate tree health index which is the ratio of healthy pixels to entire tree pixels within the tree crown. The tree was classified as healthy if the index was greater than 70%. Accuracy was checked against a random sample of 1,186 trees. At the whole tree level, 86% of campus trees were classified as healthy with 88% mapping accuracy. At the pixel level, 86% of the campus tree cover was classified as healthy. This tree health evaluation approach allows managers to identify the location of unhealthy trees for further diagnosis and treatment. It can be used to track the spread of disease and monitor seasonal or annual changes in tree health. Also, it provides tree health information that is fundamental to modeling and analysis of the environmental, social, and economic services produced by urban forests.     

Effects of ground surface permeability on the growth of urban linden trees

Street trees are an important part of urban vegetation due to their provisioning of different types of ecosystem services such as local climate regulation and contribution to aesthetical and recreational values. In order to provide these services, urban trees need to endure many stress factors not present in natural environments, such as the widespread use of impervious surfaces in the vicinity of street trees. However, few studies have evaluated the effect of this potential stress factor on urban tree growth. The aim of this study was therefore to investigate how ground surface permeability affects stem and current-year shoot growth of linden (Tilia europaea) street trees in Gothenburg, Sweden. We found that a small fraction of permeable ground surface in the vertically projected tree crown area caused lower stem growth and strongly suppressed current-year shoot growth. This finding can guide future city planning, demonstrating that the vitality of street trees is compromised when the permeable surface area in the vicinity of the tree is small.     

Structure and composition of woody plants in urban forest remnants with different adjacent land-use and slope aspect

The plant community in the karst forest is influenced by mesoclimatic conditions, such as temperature and humidity and these conditions are in turn influenced by topographic features. Urban areas adjacent to karstic forest remnants also have the potential to influence mesoclimatic variables and may serve as a source of exotic species. We described the temperature and relative humidity of karstic forest remnants at slopes with combinations of either southeast (SE) or northwest (NW) aspect, and adjacent to either urban (URB) or secondary forest (SF). We also compared forest structure and composition on these slopes to understand the interaction between forest aspect and adjacent land use on the plant community. URB-SE slopes had highest temperature and lowest humidity among all adjacent land use-aspect combinations. Understory species in SE slopes had similar richness and diversity independently of adjacent land use, however within NW aspect, slopes adjacent to urban areas showed higher values than slopes adjacent secondary forest. Canopy trees species in slopes adjacent to secondary forest were more diverse and rich in SE than in NW, although no differences between aspects were detected within URB-adjacent slopes. Understory species composition was different among the four land use-aspect combinations. However, composition of canopy trees species was different only between aspect types. Most exotic species in the sites were associated with previous agricultural practices and not with current land use. Our results show a temporal and spatial influence of anthropogenic systems on the plant community in karstic forest remnants at a suburban area of Puerto Rico.     

Mortality and recruitment in an urban forest (Forest Park in Portland,Oregon) between 1993 and 2003

I examined tree recruitment and mortality over a ten-year period at permanent plots in an urban forest, Forest Park, in Portland, Oregon. The density and diameter at breast height (dbh) for all trees living and dead were measured in 1993 and again in 2003. Data were analyzed using paired Student t-tests. I found significantly fewer live and significantly more dead trees in 2003 than in 1993. The increase in mortality was significant for all species of trees and for all sizes except large diameter trees. Mortality rates ranged from 0% to 67% at my sites. Recruitment was lower at all sites in 2003 with significantly fewer seedlings and saplings. The high mortality and low recruitment resulted in a net loss of trees at all sites. Loss of trees was not offset by increasing tree diameter, which suggests self-thinning is not the cause. No strong correlation with an urban to rural land use gradient was observed. The results may be related to global climate change or pollution. The high mortality of trees of all species in many diameter classes without a concomitant increase in recruitment could lead to dramatic changes in forest structure.     

People or place? An exploration of social and ecological drivers of urban forest species composition

Urban forests have garnered increasing attention as providers of an array of beneficial ecosystem services. However, urban forest ecosystems are highly complex and heterogeneous systems whose structure are shaped by interacting social and ecological processes. Approaches to reliably identify and differentiate these processes could be valuable for addressing complexity and reducing uncertainty in decision-making in urban forestry. The purpose of this study is to identify and quantify a range of social and ecological drivers of urban forest species composition, distribution, and diversity. This was done using hierarchical cluster analysis and discriminant analysis with empirical plot data describing the tree species composition in Toronto, Canada. Tree density and imperviousness were by far the most influential drivers of species composition. Increasing imperviousness saw not just reduced tree density but a decline in native species abundance. Additionally, single-detached housing, homeownership, and income were closely associated and explained higher tree densities and abundances of native species. However, income had a lower than expected influence on urban forest species composition given its importance in canopy cover research. Continuous forest patches were highly distinct compared to the remainder of the urban landscape, which highlights the ecological distinctiveness of residual forests within cities and lends support to their conservation. Increasing the understanding of social and ecological drivers of tree species composition, distribution, and diversity within cities is an integral part of urban forest ecosystem classification, which can be a valuable decision-support tool for ecosystem-based management in urban forestry.     

Homestead greening is widespread amongst the urban poor in three medium-sized South African towns

Trees in urban areas are important sources of ecosystem services and benefits. In most towns the bulk of urban biodiversity, and trees specifically, are found in homestead gardens. But there is only limited understanding of the tree holdings in such gardens, and how they vary within and between towns, especially for developing countries where rapid urbanisation and high poverty influence the use of and reliance on land and local resources. We report on the nature of tree holdings in private gardens of poorer suburbs in three medium-sized towns along a gradient of decreasing mean annual rainfall in northern South Africa. A total of 3 217 trees were enumerated across 450 randomly selected homesteads. Most (90 %) households had at least one tree on their homestead, with an average of 7.7?±?6.1 trees. Most householders had planted the trees themselves. The density of trees declined along the moisture gradient. Within towns, tree density was positively related to garden size, which in turn was related to relative affluence and age of the suburb. Newer and poorer suburbs had the fewest trees per household. Sixty-two tree species were recorded, which were dominated by alien species, especially fruit trees. There was no relationship between the moisture gradient and tree species richness per household, but within towns there was a difference between suburbs, being lowest in the newest suburbs. Numbers of trees and species per household was positively related to age of the household head.     

Canopy precipitation interception in urban forests in relation to stand structure

Urban forests provide important ecosystem services. In terms of hydrological benefits, forest ecosystems in urban environments represent qualitative and quantitative filter for rainwater. We quantified the canopy interception in relation to urban forest stand structure and rainfall intensity in an urban transect of the mixed (upland) forest in the city centre, towards a riparian pine forest and a floodplain hardwood forest in the City of Ljubljana, Slovenia. Bulk precipitation in open areas and throughfall were measured with fixed rainfall collectors in each forest. Stemflow was estimated from a review of relevant literature. We found that canopy interception in selected urban forests was mainly affected by tree species composition and other stand structure variables, such as canopy cover and tree dimensions. Average annual canopy interception was highest in the mixed forest (18.0% of bulk precipitation), while the riparian pine forest had the lowest level (3.9% of bulk precipitation) and the floodplain hardwood forest had the intermediate level for interception (7.1% of bulk precipitation). The mixed forest exhibited the stand structure factors that contributed to the highest canopy interception among the studied forests: high assemblage of dominant coniferous trees, denser canopy cover and the highest growing stock. Furthermore, rainfall intensity has proven to be an important factor for the seasonal partitioning (comparing the leafed and leafless period) of canopy interception. A better understanding of precipitation interception processes in urban forests is needed to assist urban forest managing and planning, aiming at maximizing canopy interception for the mitigation of stormwater runoff and flooding in urbanized watershed.     

Forest structure and anthropogenic disturbances regulate plant invasion in urban forests

Urban forest ecosystems, the structure, and functions therein are subjected to anthropogenic disturbances. Native and sensitive species from those forests might be lost due to such disturbances. At the same time, supplemented anthropogenic resources might create opportunities for exotic and invasive species. Although invasive species are considered one of the major threats to the urban biodiversity and ecosystems, the research on invasion dynamics in the Himalayas has primarily focused on the impacts of invasion on forest structure and productivity. This study aims to understand the influence of forest structure and anthropogenic factors in invasion success that are poorly covered in the existing literature. We selected 11 urban forest patches for the study considering the presence-absence of selected invasive species and structural attributes. We used Principal Component Analysis (PCA) to reduce co-linearity in the covariates and generalized linear mixed effects model (GLMM) to identify the factors affecting invasion success. We found that forest structural attributes, namely, tree diameter, height and canopy cover, and anthropogenic disturbances regulate invasion success in urban forests. This implies that maintaining urban forest structural attributes, especially the stands with large-sized trees, is essential to control invasion in the context of urbanization.

     

Woody vegetation and canopy fragmentation along a forest-to-urban gradient

To identify patterns that can be used to predict vegetation and landscape characteristics in urban environments, we surveyed the species composition and size of woody plants, as well as the landscape structure of forest canopies, along a forest-to-urban gradient near Oxford, Ohio, USA. The gradient included six sites of increasingly urban land-use: a preserve, a recreational area, a golf course, a residential subdivision, apartment complexes, and a business district. We recorded species identity and stem diameter for all woody plants greater than 3 cm diameter at breast height (DBH) to examine the distribution of individual species as well as overall community composition. We used digitized aerial photographs to compare the spatial characteristics of the forest canopy at each site. We found predictable patterns in species diversity (Shannon index), spatial heterogeneity in species composition (mean percent dissimilarity), and all measures of patch fragmentation (canopy cover and patch number and size). There were clear differences in tree density and total basal area between forested sites and developed sites, but there was little resolution among developed sites. Species richness and average DBH showed no clear pattern, suggesting that landscaping preference largely determined these values. We present a modified version of an intermediate heterogeneity model that can be used to predict diversity patterns in urban areas. We discuss probable mechanisms that led to these patterns and the potential implications for animal communities in urban environments.     

The influence of urbanization on forest stand dynamics in Northwestern Oregon

Urbanization has been shown to affect forest stand characteristics in nearby natural areas. The purpose of this study was to examine forest structure in a naturally forested area, Forest Park in Portland, OR and adjacent forested land. Tree community structure was examined at 25 sites distributed along an urban-rural land use gradient. All trees in three quadrats per site were identified to species and the diameter at breast height was measured. ANOVA was used to examine differences in species richness and diversity, and tree density and importance value among four categorical areas. Tree species richness and diversity, and the density, diameter and importance values of shade-tolerant (later successional) species of trees decreased with urbanization. Sites nearer the city of Portland had significantly fewer shade tolerant saplings and young trees and were dominated by earlier successional species of trees as compared with sites at the far end of the study area. The forest structure in the city section of the park was very similar to that in the significantly younger middle section. The lack of young shade tolerant saplings and young trees appears to be the result of urbanization, although the mechanisms for such a loss are unknown at this time. Such a lack of recruitment may interfere with normal successional processes at more urban areas of the park.     

How do people perceive urban trees? Assessing likes and dislikes in relation to the trees of a city

Cities are systems that include natural and human-created components. When a city grows without proper planning, it tends to have low environmental quality. If improving environmental quality is intended, people’s opinion should be taken into account for a better acceptance of urban management decisions. In this study, we assessed people’s perception of trees by conducting a survey with a controlled sample of citizens from the city of Morelia (west-central Mexico). Citizens liked both native and exotic tree species and rejected mainly exotic ones. Preference for trees were related to tree attributes; such as size. Trees that dropped leaves or tended to fall were not liked. The most-mentioned tree-related benefits were oxygen supply and shade; the most mentioned tree-related damages were accidents and infrastructure damage. The majority of respondents preferred trees near houses to increase tree density. Also, most respondents preferred trees in green areas as well as close to their houses, as they consider that trees provide oxygen. The majority of the respondents thought more trees were needed in the city. In general, our results show that although people perceive that trees in urban areas can cause damages, they often show more interest for the benefits related to trees and consider there should be more trees in cities. We strongly suggest the development of studies that broaden our knowledge of citizen preferences in relation to urban vegetation, and that further policy making takes their perception into account when considering creating new urban green areas, regardless of their type or size.     

Predicting tree species richness in urban forests

There has been an increasing interest in urban forests and the levels of biodiversity they contain. Currently there are no spatially explicit maps of tree species richness in urban areas. This research tests and identifies GIS and remote sensing metrics (climate, area, productivity, three-dimensional structure) hypothesized to be associated with species richness in native forests and identifies methods that can be applied to predict and map tree species richness in cities. We quantified tree species richness, floristic composition, and structure in 28 1-ha plots in the city of Los Angeles. Climate and remote sensing metrics from high-resolution aerial imagery (10 cm), QuickBird (60 cm), Landsat (30 m), MODIS (250 m), and airborne lidar (2 m) were collected for each plot. There were 1208 individual stems and 108 trees identified to species. Species richness ranged from 2 to 31 species per ha and averaged 17 species per ha. Tree canopy cover from QuickBird explained the highest portion of variance (54%) in tree species richness followed by NDVI from Landsat (42%). Tree species richness can be higher in residential urban forests than native forests in the United States. Spatially explicit species richness maps at 1 ha can be created and tested for cities in order to identify both hotspots and coldspots of tree species richness and changes in species richness over time.     

Rainfall interception by Santa Monica's municipal urban forest

Effects of urban forests on rainfall interception and runoff reduction have been conceptualized, but not well quantified. In this study rainfall interception by street and park trees in Santa Monica, California is simulated. A mass and energy balance rainfall interception model is used to simulate rainfall interception processes (e.g., gross precipitation, free throughfall, canopy drip, stemflow, and evaporation). Annual rainfall interception by the 29,299 street and park trees was 193,168 m³ (6.6 m³/tree), or 1.6% of total precipitation. The annual value of avoided stormwater treatment and flood control costs associated with reduced runoff was $110,890 ($3.60/tree). Interception rate varied with tree species and sizes. Rainfall interception ranged from 15.3% (0.8 m³/tree) for a small Jacaranda mimosifolia (3.5 cm diameter at breast height) to 66.5% (20.8 m³/tree) for a mature Tristania conferta (38.1 cm). In a 25-year storm, interception by all street and park trees was 12,139.5 m³ (0.4%), each tree yielding $0.60 (0.4 m³/tree) in avoided flood control costs. Rainfall interception varied seasonally, averaging 14.8% during a 21.7 mm winter storm and 79.5% during a 20.3 mm summer storm for a large, deciduous Platanus acerifolia tree. Effects of differences in temporal precipitation patterns, tree population traits, and pruning practices on interception in Santa Monica, Modesto, and Sacramento, California are described.