Tree diversity,distribution, history and change in urban parks: studies in Bangalore,India

Urban parks constitute critical biodiversity hotspots in crowded, concrete-dominated city environments. Despite the importance, they remain little researched. This paper assesses the biodiversity and distribution of trees in urban parks in the southern Indian city of Bangalore. 127 plots were used to survey tree distribution in parks across the city. The distribution is largely dominated by a few common species. The proportion of exotic species was very high, with 77% of trees belonging to introduced species. Park history had an impact on distribution. Old parks had fewer but larger trees, and greater species diversity compared to recently established parks. Old parks also differed in species composition, having a greater proportion of large canopy trees compared to young parks. Examination of size distributions revealed that large canopied species were gradually being phased out, and replaced by narrow and medium sized tree species which are easier to maintain, but which may not provide the same environmental and ecological benefits. Greater attention requires to be paid to the selection of trees in cities, not just with a view to easy maintenance as is currently the case, but to select an appropriate mix of trees that supports biodiversity and maximizes environmental and ecosystem services.     

Effects of urbanization on butterfly species richness,guild structure,and rarity

We evaluated the effects of landscape characteristics associated with urbanization, as well as local features, on butterfly species richness at four spatial scales (50, 150, 500, and 1,000 m from survey plots). We also evaluated these effects separately by butterfly guilds based on their region-wide rarity and on degree of specialization. The distribution of abundances of the 44 species observed showed an excess of uncommon species compared to a log-normal distribution, and the two most abundant species were exotic (Thymelicus lineola and Pieris rapae). We used an information theoretic approach to model selection to determine the most important correlates of butterfly species richness. Models of mean butterfly richness per visit explained greater variance than did models of cumulative richness across the season. Cumulative butterfly species richness was affected more at larger spatial scales, while richness per visit was affected similarly at all spatial scales. The most consistent local factor affecting butterfly diversity was the number of nectar producing plants that were flowering. The most consistent landscape-level correlates of species richness were number of people (at small spatial scales) and green space. Measures of increased urbanization were associated with decreased butterfly species richness, and rare and specialized species were most affected. Species that were regionally rare, and those that specialized both in host plants and had few broods, disappeared with declining richness across sites 2.9–4.5 times faster than did generalists and less restricted specialists.     

Native and exotic species in the urban landscape of the city of Rio de Janeiro,Brazil: density,richness, and arboreal deficit

Shaped by European influences, the Brazilian urban landscape was marked by the predominant use of exotic species in planted areas. In the 19th century along the city streets of Rio de Janeiro, trees were planted at a standard distance from each other. With time, native species began to be used in the urban landscape. Our purpose was to evaluate the utilization of exotic plant species in the urban landscape of Rio de Janeiro, evaluate the city’s arboreal deficit, and relate its neighborhood arboreal density to its economic index. Arboreal deficit represents the negative difference between the total number of expected trees and the observed number in the streets based in the standard distance used in the past. Twenty native and 40 exotic species were found in the 1701 streets sampled. A high percent of streets did not have any planted trees and the number of trees was greater in wealthier neighborhoods. The strong prevalence of exotic species is indicative of the negative impact of human activity on the biota. Since the municipality of Rio de Janeiro harbors forested areas, the risk for dissemination of exotic species is high.     

The effects of trampling on assemblages of ground beetles (Coleoptera,Carabidae) in urban forests in Helsinki,Finland

The occurrence of carabid beetles in relation to trampling was examined in urban forest sites located in the city of Helsinki, southern Finland. The degree of wear of the forest floor was assessed and used as a measure of trampling intensity. In particular, we examined the following predictions: (1) carabid diversity should decrease with increasing trampling intensity, (2) mean body size of the dominating carabid species should decrease with increasing trampling intensity, and (3) opportunistic species should gain dominance in severely trampled sites. In total, 1,326 beetles representing 27 species were captured. The first prediction was not supported, as there was no correlation between species richness or Hill's N2 diversity index and trampling intensity. However, there was a positive correlation between number of carabids captured and trampling intensity of the site. The second hypothesis gained some support, as there was a marginally significant negative correlation between body size and trampling intensity. The hypothesis that opportunistic species should gain dominance in severely trampled sites was supported as one species was very dominant (Pterostichus melanarius, 60.0%) in the heavily trampled sites, while there were two equally and less dominant species in the less trampled sites. Individual species did show different responses to the effects of trampling, and the most sensitive forest species may not survive in the heavily trampled sites. We conclude that at the community level (e.g., species richness, diversity), the effects of trampling on carabids in urban forests are subtle, but impacts are pronounced for some sensitive species.     

Interactions between birds, fruit bats and exotic plants in urban Hong Kong, South China

I observed the use of exotic plant species by native birds and bats in 8 km² of urban Hong Kong. Twenty-nine plant taxa were used repeatedly. Five species accounted for most interactions with birds: Cinnamomum camphora (10 bird species), Ficus religiosa (6), Livistona chinensis (7) and Pyracantha crenulata (6) provided fruits, while Bombax ceiba (6) supplied nectar. Panicum maximum (2) and C. camphora (1) provided seeds for granivorous birds and Leucaena leucocephala is host to a psyllid consumed by small insectivores in winter. Most trophic interactions between exotics and fruit bats involved the fruits or flowers of Syzygium jambos, while Livistona chinensis provided roosting sites for Cynopterus sphinx. The fruiting phenologies of the exotic species differed from the local flora, with fruit of some species available for the majority of the year. Fruit characteristics for the most commonly eaten species are within the range for native fruit species, although most are larger and have a lower seed load than the medians for native species. Seed dispersal by birds and/or fruit bats is a necessary, but not sufficient, condition for the naturalization of fleshy-fruited plants in Hong Kong, but S. jambos is the only naturalizing exotic species that needs vertebrates for pollination.     

Effects of area size,heterogeneity, isolation,and disturbances on urban park avifauna in a highly populated tropical city

We assessed bird diversity and nesting activity in 54 urban parks in the core of a highly populated East-Asian tropical city to investigate the effects of area, isolation, habitat heterogeneity, and human disturbances on avifauna. Tree density and heterogeneity were lower while isolation was greater in small than in large or medium-sized parks. Yet, park features were often interrelated, with isolation variables negatively correlated to each other, heterogeneity positively correlated to area, and habitat variables mostly positively correlated with one another but variously to disturbances; whereas pedestrian densities wee positively correlated with green area proportions (ISO3) but negatively with the distances to large green areas (ISO2). Park size played a primary role by positively affecting species richness and numbers of nesting species, thus contributing to higher total and nesting species heterogeneity, and determining the occurrence of a greatest number of species. Pedestrian density, canopy cover, and ground heterogeneity played secondary roles, whereas isolation appeared less influential at the community level. Omnivores, granivores, and aerial insectivores appeared more abundant but were dominated by several common species. Nesting was restricted to even fewer species. While species varied in the occupancy responses to these factors, tree heterogeneity positively affected the nesting of Zosterops japonicus and cavity-nesting birds, and predator density negatively affected those of Lonchura punctulata and Streptopelia doves. Improved urban land use planning and park management incorporating these effects should be implemented to enhance breeding and avoid compositional homogenization for long-term sustainability of urban avian diversity.     

Woody plant communities along urban,suburban, and rural streams in Louisville,Kentucky, USA

Anthropogenic changes in land use and cover (LULC) in stream catchments can alter the composition of riparian plant communities, which can affect ecosystem functions of riparian areas and streams from local to landscape scales. We conducted a study to determine if woody plant species composition and abundance along headwater streams were correlated with categorical and continuous LULC and environmental variables along an urban-to-rural gradient. These variables were calculated at different spatial scales (subcatchment level and within 0.5 and 1 km radii of plots) and used % impervious surface cover (ISC) and finer scale LULC classification levels to determine their ability to explain species composition, diversity, abundance, non-native provenance and wetland indicator status of four plant strata—canopy tree, tree sapling, tree seedling and shrub. At all scales, we found distinct riparian woody communities within land-use categories, with significant differences among shrub species. Fine-scale land-cover variables correlated with species composition of shrub, tree sapling and tree seedling strata, but not the canopy tree stratum. Celtis occidentalis and Acer negundo were ubiquitous but dominated riparian areas surrounded by development, while Asimina triloba was associated with forested rural riparian banks. Non-native shrubs, Lonicera maackii and Euonymus alatus, were indicative of areas surrounded by development, while the native shrub, Lindera benzoin, was associated with deciduous forest. Negative factor-ceiling relationships between canopy tree, sapling and tree seedling densities and % ISC were found, with abrupt declines above approximately 30 % ISC. Facultative wetland shrubs were not found above 30 % ISC. Streambank height, which was strongly negatively correlated with depth to the water table and positively correlated with cumulative catchment area, was negatively correlated with facultative wetland tree and shrub species. In addition, riparian tree sapling and seedling densities declined as the abundance of Lonicera maackii increased.     

Rape in nonhuman animal species: Definitions,evidence, and implications

This paper examines the controversial issue of rape in nonhuman animal species. Many existing studies on nonhuman rape appear to suffer from inappropriate definitions and questionable interpretations. Many existing critiques of these studies are limited by their own misunderstandings of evolutionary theory. An attempt is made to reduce the resulting controversy by defining rape in a way that can be applied to nonhuman animals, without provoking unwarranted implications about human rape. Applying this definition to existing studies indicates that behavior meeting the definition of rape does exist in several nonhuman species. However, comparisons of these species do not provide clear implications for either proximate or ultimate explanations of human rape.     

An investigation of the composition of the urban riparian soil propagule bank along the River Brent, Greater London, UK, in comparison with previous propagule bank studies in rural areas

Most propagule bank research has been conducted in rural areas and has emphasised agricultural soils, although research on rural riparian propagule banks has expanded rapidly over the last decade. This paper presents observations of the species abundance of plant propagules in urban riparian soils along the River Brent, Greater London, UK. The data are analysed to explore changes in species abundance of the propagule bank with depth (0?C5?cm compared with 5?C10?cm soil layer), with distance from the river (0?C1?m, 1?C2?m, 2?C3?m), and at different locations on the river network. No significant differences were found in propagule abundance with soil depth or distance from the river??s edge, but significantly more species were found in the propagule bank close to the river??s edge. The species composition of the Brent??s propagule bank is then compared with riparian propagule banks from the rural rivers Dove, Frome and Tern, UK. Species richness of the Brent propagule bank samples was generally lower than was found in the three rural rivers and, in total, 21%, 4%, 5% and 4% of observed species were aliens in the Brent, Dove, Frome and Tern soil samples, respectively. When the species present in the Brent??s propagule bank were compared with published data on urban garden soils in another UK city, the latter were found to support a higher number of alien species, suggesting that other propagule sources to urban river systems support lower numbers of alien species in riparian zones than domestic gardens.     

Bird species assemblages across a rural urban gradient around Kolkata,India

Urban development leads to changes in habitat structure and resource base. Bird communities are known to respond sharply to such changes. Our result from study of bird community along urbanization gradient around Kolkata metropolitan city clearly separated the urban bird community from the rural and rural 3 habitats in terms of species diversity and foraging groups. Rural and rural 3 sites had more number of rare species and higher percentage of unique species as compared to the urban habitats. Functional group analysis showed higher abundance of granivores in urban habitats and absence of insectivore and carnivorous species that were found in the rural and rural 3 habitats. The bird species assemblage along the gradient was significantly nested where bird species recorded in urban areas were subset of the species rich rural areas. There was no difference in individual counts between urban and rural habitats, therefore nullifying the hypothesis that rural areas are more species rich because of higher population size. Bird community in the urban areas was less even as compared to the rural areas due to the dominance of omnivorous guild. Bird diversity was negatively correlated to the density of house lots.     

Characteristics of the nonelderly with selected sources of health insurance and lengths of uninsured spells

This Issue Brief examines the characteristics of individuals with selected sources of coverage and combinations of sources of coverage over a 12-month period. In addition, it examines the characteristics of individuals who experience spells without health insurance and the lengths of these spells. It uses the most recent 12-month period from the Survey of Income and Program Participation and builds on previous research on the lengths of spells with and without health insurance. Analysis of individuals' health insurance coverage from October 1994 to September 1995 showed that approximately 77.6 percent of the nonelderly had health insurance coverage during this entire period. In addition, 22.4 percent of the nonelderly were uninsured for at least one month during this period, and 7.4 percent of the nonelderly were uninsured for the entire period. Of those with health insurance coverage for the entire year, approximately 83 percent were covered by private health insurance, with at least 81 percent of this group receiving the coverage from employment-based sources. Eighty-five percent of the spells without health insurance with an observed beginning and end lasted for 4 months or less, and 99 percent lasted for 8 months or less. When examining the spells with either an observed beginning or end, 55 percent of these spells were found to last for 4 months or less, and 87 percent were found to last for 8 months or less. However, investigation of all spells without health insurance showed that approximately one-half of all spells without health insurance coverage lasted for 8 months or longer. This report found that two-thirds of spells without health insurance last for less than one year, confirming previous research that a majority of these spells are for less than a year. However, this report also confirms the existence of a significant number--approximately one-third of all individuals with a spell of noncoverage--of chronically uninsured individuals. These individuals are the most likely to delay seeking treatment for illnesses and to use the emergency room as their only site of care. Because they are in poverty or near poverty, much of this care is uncompensated. Thus, to the extent that providers can shift these costs onto other payers, all individuals and employers share in these costs through higher health insurance premiums or higher taxes to finance public hospitals and public insurance programs. Recent major health insurance legislation has addressed access to health insurance, and in many cases focused solely on continued access to employment-based coverage, but has done very little to address the affordability of coverage. However, as this report demonstrates, many individuals experiencing spells without health insurance have low incomes. Thus, to obtain coverage, individuals need not only increased access to health insurance but also the ability to afford this health insurance.     

Habitat type,food resources,noise and light pollution explain the species composition,abundance and stability of a winter bird assemblage in an urban environment

At present, urban areas cover almost 3% of the Earth’s terrestrial area, and this proportion is constantly increasing. Although urbanization leads to a decline in biodiversity, at the same time it creates extensive habitats that are exploited by an assemblage of organisms, including birds. The species composition and density of birds nesting in towns and cities are determined by the types of buildings, the structure and maturity of urban greenery, and habitat diversity. In contrast, the habitat traits shaping the community of birds wintering in urban areas are not known. The aim of this work was to assess the influence of habitat structure, food resources and the urban effects (pollution, noise, artificial light) on an assemblage of birds overwintering in an urban area. It was carried out in 2014 and 2015 in the city of Kraków (southern Poland), on 56 randomly chosen sample plots, in which the composition, density and interseasonal similarity of bird assemblage were assessed with line transect method. A total of 64 bird species (mean = 17.7 ± 4.9 SD species/plot) was recorded. The mean density was 89.6 ind./km ±63.3 SD. The most numerous species were Great Tit Parus major, Magpie Pica pica, Blackbird Turdus merula, Blue Tit Cyanistes caeruleus, Rook Corvus frugilegus, Fieldfare Turdus pilaris and House Sparrow Passer domesticus. Noise adversely affected species numbers and density, but artificial light acted positively on the density of birds and their interseasonal stability. The species richness and density of birds were also determined by the number of food sources available (e.g. bird-feeders). In addition, the greater the proportion of open areas, the fewer species were recorded. In contrast, the more urban greenery there was, the greater the density of the entire bird assemblage. Urban infrastructure (buildings, roads, refuse tips) had a positive effect on the interseasonal stabilization of the species composition of wintering birds. The results of this work indicate that the urban effect, i.e. noise and light pollution, apart from purely habitat factors, provide a good explanation for the species richness, density and stability of bird assemblage wintering in urban areas.     

Analysis of breeding bird communities along an urban-rural gradient in Berlin,Germany, by Hasse Diagram Technique

Breeding bird communities in Berlin were investigated at five study sites, which represent an structural urban-rural gradient, from densely built-up areas with sealed ground to green areas such as large parks. In this paper, we tested the Hasse Diagram Technique (HDT), an approach belonging to partial order theory, for its appropriateness to extract information about changes in species composition from our data set. As expected, the overall number of bird species was positively correlated with the urban-rural gradient, increasing from 12 found in the city center to 28 in an urban park. By HDT analysis, four different sub-groups of birds could be identified, contributing differentially to the increase in species number: (1) ubiquists, i.e., species found at each study site, (2) a sub-group, the species’ number of which was positively correlated with the urban-rural gradient, increasing with increasingly green areas, (3) a sub-group, the species’ number of which was negatively correlated with the urban-rural gradient, decreasing with a declining density of buildings, and (4) species, the distribution of which did not show any correlation with the urban-rural gradient. The results show that HDT is a suitable tool for formalized analysis of the pattern of species’ communities along an environmental gradient.     

Bird community responses to different urban conditions in La Paz,Bolivia

Urbanization alters ecosystems worldwide, but little is known about its effects in the Neotropical region. In the present research we examined the relative influence of different levels of urbanization and of some urban development measures on bird species richness, abundance and composition. We surveyed 104 observation stations at which we collected data on the relative abundance of bird species, and also data on seven environmental variables as measures of urban development and human activity. We registered 57 native bird species. Bird species richness and bird abundance increased with lower urbanization levels. Both variables were positively related to vegetation cover and native vegetation, and negatively to built-up cover, abundance of Rock Pigeon (Columa livia), pedestrian rate and car rate. A canonical correspondence analysis produced a significant model that explained 37% of the total variation in species data. This analysis segregated bird species along two important gradients: urbanization and elevation. The most urbanized areas were dominated by a few synanthropic species tolerant to human disturbance, such as Rufous-Collared Sparrow (Zonotrichia capensis), Eared Dove (Zenaida auriculata) or Chiguanco Thrush (Turdus chiguanco). Areas with lower urbanization levels had more species typically associated with native vegetation, such as Plain-Mantled Tit-Spinetail (Leptasthenura aegitaloides) or D’Orbigny’s Chat-Tyrant (Ochthoeca oenanthoides), among others. Elevation had a significant influence in structuring bird communities, with some species restricted to higher elevations and some to lower elevations. Although changes in elevation had an important influence, urbanization had a stronger effect on structuring bird communities. This study provides valuable information and an important baseline for future studies.     

Diversity,abundance, and species composition of ants in urban green spaces

Urbanization threatens biodiversity, yet the number and scope of studies on urban arthropod biodiversity are relatively limited. We sampled ant communities in three urban habitats (forest remnants, community gardens, vacant lots) in Detroit and Toledo, USA, to compare species richness, abundance, and species composition. We measured 24 site characteristics to examine relationships between richness and composition and habitat patch size, vegetation, and urban features. Ant richness was higher in forests (26) than in gardens (14) and intermediate in vacant lots (20). Ant richness in gardens and vacant lots negatively correlated with abundance of an exotic ant species (Tetramorium caespitum); thus this ant may affect native ant richness in urban habitats. Ant composition differed with habitat type, and abundance was lowest in forests. Site characteristics varied with habitat type: forests were larger, had more woody plants, higher woody plant richness, more branches, and leaf litter whereas lots and gardens had more concrete and buildings. Vacant lots had taller herbaceous vegetation, and gardens had higher forb richness, density, and more bare ground. Differences in vegetation did not correlate with ant richness, but several vegetation factors (e.g. patch size, number and size of trees, leaf litter, and amount of concrete and buildings) correlated with differences in ant species composition. Additional factors relating to soil, nests, or microclimatic factors may also be important for urban ant communities. Implications for biodiversity conservation in urban ecosystems are discussed.     

Effects of park size,peri-urban forest spillover,and environmental filtering on diversity,structure, and morphology of ant assemblages in urban park

Urban Ecosystems - Urban parks are likely the last green areas to preserve fauna diversity in urban ecosystems. We predicted that 1) species richness and ant diversity would increase with...     

The seed bank of a restored tidal freshwater marsh in Washington,DC

The species composition of seed banks may be a useful indicator of vegetation structure and dynamics, but seed bank studies have rarely been applied to monitoring of wetland restoration projects. We studied the seed banks and vegetation of three distinct sites within a restored tidal freshwater marsh and of two nearby natural marsh sites. Surface soil samples were collected and subjected to flooded and nonflooded conditions in a greenhouse for enumeration of emerging seedlings. Percent coverage of vegetation was estimated in permanent 1-m² plots at each soil sampling location.Species composition of the seed bank and vegetation differed considerably; only 43% of the species observed were common to both. The seed bank of restored marsh sites had higher species richness and seed density than the natural marsh sites, but both had similar species diversity. Annual species were more important in the seed bank and less important in the vegetation of restored sites than reference sites. Flooding significantly reduced the species richness, diversity, and density of emerging seedlings. Most species planted at the site were rare in the seed bank, with Leersia oryzoides being an important exception. Other species abundant in the seed bank included Cyperus odoratus, Juncus effusus, and the nonindigenous Lythrum salicaria. A possible mechanism explaining the higher densities of seeds in restored areas is that ruderal vegetation colonized the restoration site, reproduced, and set seed shortly following placement of dredge material. Perennial species subsequently colonized and were planted, and expanded to reduce the abundance of annuals in restored marsh vegetation. Our study suggests that the plant regeneration dynamics of the restored marsh sites are considerably different from those of natural marshes. Additionally, we suggest that seed bank studies are a useful technique for assessing the community structure and vegetation dynamics of restored wetlands.     

Species richness,phylogenetic and functional structure of bird communities in Chinese university campuses are associated with divergent variables

University campus is an important component of urban landscapes for biodiversity conservation. However, to our knowledge no study has quantitatively assessed the diversity and structure of bird communities in Chinese university campuses, especially from phylogenetic and functional perspectives. Here, for the first time we linked species richness, phylogenetic structure and body mass structure of campus bird communities with contemporary climate, glacial-interglacial climate change, altitudinal range, population density around campus, area and age of campus to test their associations. We found 393 bird species in 38 university campuses (29% of all Chinese bird species, two species are endangered, four species are vulnerable, and 33 species are near threatened). The variables significantly correlated with campus bird species richness, phylogenetic structure and body mass structure were altitudinal range and mean annual precipitation, glacial-interglacial anomaly in temperature, and altitudinal range, respectively. In particular, there were more species in steeper and wetter campuses, more young species clustered in campuses with stable glacial-interglacial climate, and more species with smaller body size in steeper campuses. Our study highlights the importance of considering both phylogenetic and functional information for biodiversity conservation in urban ecosystems.     

Relative importance of habitat quantity, structure, and spatial pattern to birds in urbanizing environments

Urbanization reduces the quantity of native vegetation and alters its local structure and regional spatial pattern. These changes cause local extirpations of bird species associated with native vegetation and increases in the abundance and number of bird species associated with human activity. We used 54–1 km² landscapes in the Seattle, Washington, USA metropolitan area to determine (1) the relative importance of habitat quantity, structure, and pattern to bird diversity and abundance and (2) whether housing developments can be managed to mitigate the negative impacts of urbanization on forest bird diversity. In general, bird species richness was high and many native forest species were retained where urban landcover comprised less than 52% of the landscape, tree density (especially that of evergreens) remained at least 9.8 trees/ha in developments, and forest was at least 64% aggregated across the landscape. These results suggest that the quantity, structure, and pattern of forested habitat affected breeding bird diversity in urbanizing landscapes. However, habitat pattern appeared less influential than other habitat attributes when results from all community- and population-level analyses were considered. Conservation of native birds in reserves can be supplemented by managing the amount, composition, structural complexity, and—to a lesser extent—arrangement of vegetation in neighborhoods.     

Patterns of plant species diversity in deciduous woodlands of Kyiv,Ukraine

Urban Ecosystems - The present study is aimed at exploring native and alien plant species diversity patterns in distinct types of the urban forest environment by documenting differences in species...