Classes of protection in urban forest fragments are effectiveless in structuring butterfly assemblages: landscape and forest structure are far better predictors

Urbanization is a major cause of biodiversity loss and disconnection between people and nature. For this reason, the creation and maintenance of green areas as part of the urban landscape is a common practice in cities around the world. These green areas are generally a result of particular public policies regarding how these areas can be accessed, maintained, and how they are connected to other natural fragments in the city. In an effort to investigate aspects of the ecology of such green areas of the city of Curitiba, southern Brazil, the present study measured distinct parameters of the structure of butterfly assemblages from 19 forest fragments of distinct classes of protection, testing the influence of different policies (protection category) on butterfly assemblages. Comparatively, we also tested the predictive power of landscape variables (at distinct scales) and fragment quality (e.g. diameter at breast height, density of vegetation) using Generalized Linear Models (GLMs). Overall, 14,596 individuals belonging to 412 butterfly species were recorded, but values of species diversity and composition are very heterogeneous, indicating that the protection categories do not significantly interfere with the structure of the butterfly assemblages. More important than the protection category is the forest fragment quality and landscape connectivity. Our models showed that degraded fragments tend to present a specific butterfly composition, whereas the more urbanized landscape have poor butterfly species richness. These results reinforce the importance of implementing public policies that prioritize the conservation of the quality of forest fragments in every class of protection, as well as the conservation of forest fragments throughout the urban landscape.

     

How to improve urban greenspace for woodland birds: site and local-scale determinants of bird species richness

Wooded habitats represent hotspots of urban biodiversity, however, urban development imposes pressure on biota in these refuges. Identification of the most influential habitat attributes and the role of local urban characteristics is crucial for proper decisions on management practices supporting biodiversity. We aimed to identify well manageable fine-scale habitat attributes to suggest specific, feasible and affordable management recommendations for green space in cities. We analysed species richness of woodland-associated bird communities and incidence of individual species at 290 sites in a wide variety of green areas scattered across the city of Prague, Czech Republic. Generalized linear mixed models (GLMM) and regression tree analyses were used to identify site-scale (100 m radius sampling sites) and local-scale (200 m and 500 m radius plots) habitat attributes shaping the bird communities at individual sites. Logistic regression was used to assess the responses of individual species to habitat characteristics. Our results imply that at the site scale, management practices should focus on maintenance and promoting species-diverse and older tree stands, with a mixture of coniferous and deciduous trees. Water-bodies and accompanying riparian habitats should be maintained and carefully managed to preserve high-quality remnants of natural vegetation. Presence of a few old trees (about 12 % of tree cover with DBH?>?50 cm) or small urban standing water and watercourses enrich the bird community by at least two species. Species richness of woodland avifauna at particular sites is further supported by the total amount of tree cover in the surroundings, including scattered greenery of public spaces and private gardens. We conclude that proper management at site scale has the potential to increase biodiversity of the urban environment.     

Reducing management intensity and isolation as promising tools to enhance ground-dwelling arthropod diversity in urban grasslands

Urbanisation is a growing global phenomenon having multiple ecological consequences. However, the effects of urbanisation on biodiversity remain ambiguous, and some evidence exists that cities provide valuable secondary habitats for many species, possibly mitigating regional biodiversity loss. Therefore, the value of urban sites for biodiversity depends on local habitat conditions and the configuration of the surrounding landscape. We aimed to disentangle the effects of local habitat parameters (management, plant diversity) and urban matrix variables (urbanisation, isolation) on carabid beetle and spider diversity and traits in informal urban green spaces in Berlin, Germany. Habitat management and isolation were the most important influences on carabid beetle and spider species and trait compositions. Spider communities of irregular managed sites contained 2.5 times more species of conservation concern than extensive (regular) managed sites. Moreover, irregular managed sites contained larger species (both for carabid beetles and spiders) and affected the hunting mode of spiders. Isolated sites tended to have lower spider species richness and number of spider species of conservation concern. Moreover, isolated sites were characterised by small, mobile and herbivorous carabid beetles. In contrast, urbanisation and local plant diversity had no effect on carabid beetles and spiders. We conclude that urban grasslands within residential areas – even if not targeted for conservation plans – can provide important habitats for conserving biodiversity, including species of conservation concern. Reducing the intensity of habitat management and increasing the connectivity of urban grassland sites can promote diverse arthropod communities and should therefore be considered in urban planning.     

Management of nature-based goods and services provisioning from the urban common: a pan-European perspective

The role of the urban common (i.e. shared space and resources) in sustainable provisioning of goods and services to city dwellers is discussed in this paper. Focusing on tree-based green infrastructure, the study scope includes three categories of provisioning (woody biomass, food/fibre, and non-timber forest products, i.e. NTFPs), alongside three categories of supporting services (fresh water replenishment, soil nutrient restoration, building preservation). As a first step, prospects of utilizing the urban common as facilitator of nature-based solution to the earmarked provisioning services are evaluated through dedicated literature survey and expert elicitation on perceived impact of environmental change triggers and management interventions (planning and/or governance). This is followed by a structured review of the state of affairs in four European cities (London, Amsterdam, Sofia, Ljubljana), representing different macro-geographical regions with distinct socio-economic drivers in managing these provisioning services. The pan-European expert elicitation exercise noted active management of the urban common as positively impacting on the performance of the majority of provisioning services, while environmental change impacts were found to be overriding and adversely influencing the provisioning of material resources (mainly NTFPs and woody biomass). The four-city case study highlighted some regional peculiarities in connecting the city dwellers to the urban common and identified the need to overcome socio-cultural barriers for enhancing pan-European best practice sharing in the management of goods and services provisioning. This is deemed essential to pave way for an emerging perspective on sustainable utilization of the urban common as an enabler for nature-based solution, making it fit for purpose in meeting the astronomical demands of future urban living.

     

How small cities affect the biodiversity of ground-dwelling mammals and the relevance of this knowledge in planning urban land expansion in terms of urban wildlife

Fundamental principles regarding urban biodiversity are based on studies conducted in large cities. However we cannot know whether the same phenomena occur in smaller cities or how small cities affect biodiversity. Small cities are an inherent element of urbanization and in the future, most global urban growth is expected to take place in small and medium-sized cities. Understanding the effects of small cities on biodiversity will be an important aspect in planning urban land expansion. Our study examined the effects of a small city on communities of small ground-dwelling mammals on 41 sites arranged in a four step gradient of urbanization. In 6700 trap-days, we caught 2333 individuals comprising 15 species. In the downtown area the same phenomena as those described for large cities were observed: a reduction in species richness and diversity, a decline in the abundance of urban sensitive species and an increase in synurbic species. However, in contrast to large city studies, green areas outside the downtown area did not differ from rural sites in small mammal population parameters. This phenomenon of relatively unchanged fauna outside the downtown area shows that small cities have the potential to maintain a high level of diversity of small ground-dwelling mammals if appropriate planning of further building expansion is implemented. More studies of small cities are needed to better assess their impact on biodiversity. This knowledge can then be applied in better planning for urban wildlife. Generalizations based solely on large city studies are inadequate and may lead to incomplete or inappropriate conservation strategies for small cities.     

Ecological lands for conservation of vascular plant diversity in the urban environment

Conservation of biodiversity in urban areas has become crucial to urban green area management. There are several legislative solutions for preservation of species and habitats in cities. One of them is ‘ecological lands’ – a low-restrictive form of protected areas in Poland. We aimed to assess their efficiency in vascular flora biodiversity conservation in the urban environment in Poznań (W Poland; 550,000 inhabitants). We hypothesized that ecological lands which cover <2% of the city area comprise over 50% of taxonomic diversity and over 90% of functional trait-level range of the vascular flora. Analysis of five ecological lands, which covered 1.8% of the whole city area confirmed our hypothesis. In ecological lands studied, we found 564 species of vascular plants, which is 52.9% of the whole city flora. These species belonged to 23 of 29 phytosociological classes represented in the whole city (73.9%). Functional trait distributions in ecological lands studied comprised from 95.8 to 100% of trait distributions in the flora of the whole city. Ecological lands seem to be a good way for conservation of biodiversity in urban areas. The legislative simplicity and low restrictiveness for both land management and recreational utility make ecological lands a much easier form of nature conservation which may be adapted to other cities for more efficient biodiversity management.     

Effects of biodiversity and environment-related attitude on perception of urban green space

Green space in cities contributes to the quality of life for city dwellers, e.g., by increasing the opportunity for recreation. However, perception of urban green space is influenced by multiple factors. We investigated effects of biodiversity and environment-related attitudes on visual and auditory perceptions of urban green space. Field measurements of biodiversity were conducted in six sites across an urban gradient in Gothenburg, Sweden, and three categories of biodiversity—high, medium, low—were established. Households were sent a survey on aesthetic perception of urban green space, sound perception and the importance of trees and plants for the perception of bird species. Each respondent focused on the site that was located nearby. The environment-related attitudes comprised “Nature-oriented” and “Urban-oriented” persons and were based on participants’ own attitude estimations. It was shown that participants’ “subjective” aesthetic and sound-related perception of urban greenery were in line with the “objectively” measured subdivisions of high, medium and low biodiversity. So also were their estimations of the importance of trees and plants for perception of bird species in urban greenery, although differing only between high and medium/low biodiversity conditions. Persons rating themselves as highly nature-oriented were shown to give higher scores to urban green space aesthetics and to value greenery-related sounds higher, and to attach greater importance to trees and plants in their perception of bird species in urban greenery, than less nature-oriented persons. Highly urban-oriented persons compared to less urban-oriented persons did the same, but only regarding urban greenery-related aesthetics and sounds of nature. We conclude that environment-related attitudes influence perceptions of green space. Moreover, our findings support the idea that biodiversity per se also influences perceptions; people value green space significantly more with high than with low measured biodiversity. Urban planning needs to provide city inhabitants with green spaces that are species-rich, lush, varied and rich with natural sounds.     

Evaluating the effects of canine urine on urban soil microbial communities

Due to extensive areas of impermeable surfaces, combined sewer overflow (CSO) is currently a major problem in urban areas across the United States. In CSO systems, sewage can travel through underground pipes to be decontaminated in treatment facilities, or it can combine with stormwater after a precipitation event and discharge into local waterways. Many cities are implementing green infrastructure installations, which use vegetation and bioactive soil microbial communities to enhance soil water-holding capacity, thereby minimizing CSO events. Understanding the factors that structure soil microbial communities in green infrastructure will facilitate more effective management of these engineered ecosystems; however, few studies to date have evaluated ecological patterns and processes of microbes in the urban environment. Nitrogen loading is known to be a major factor structuring fungi and bacteria in non-urban soils, and since cities also contain large populations of canines, N-rich urine deposition is a potential factor that could be important for structuring soil microbes in ground-level green infrastructure installations. Our study investigated the effects of canine urine on the urban soil microbial communities in a greenhouse experiment by treating Liriope muscari, a common plant found in New York City green infrastructure, with different concentrations of canine urine for 4 weeks in an experimental setting. We found that urine application significantly decreased total soil microbial biomass and microbial richness, and increased water runoff volume. These findings indicate that canine urine may have negative consequences for soil water-holding capacity and nutrient cycling in urban green infrastructure installations by directly decreasing the abundance and richness of soil microbial communities.

     

Relationship of bird richness,abundance and assemblage to the built environment in a small island tropical urban setting: a Suva,Fiji case study

There have been few studies investigating the relationship between the built environment and the status of bird distributions in small island tropical urban areas. We present a study investigating the relationship between bird species richness, abundance and assemblage to the built environment in Suva, Fiji. Field surveys were taken at 54 randomly selected sites throughout the city, stratified by three building density classes and the central business district (CBD). At each site bird counts were recorded, along with environmental data such as average building height, within a 150 m radius. Land-use information was obtained from screen digitized high-resolution satellite imagery within the same radius. Distance to undeveloped patches of land within the urban area was calculated using a GIS. Analysis of the effects of the built environment was carried out for all species, and for exotic and native species separately. Abundance of exotics was significantly higher in the central business district (CBD) than all other urban density classes, and significantly higher than natives in all other density classes. We found a negative relationship between native species richness and distance to undeveloped patches, but no relationship for exotics. Species assemblage was not related to urban density class. We conclude that the status of native and exotic bird species in Suva is similar to what has been found in urban areas in temperate climates, and conservation efforts should focus on minimizing the amount of heavily urbanized “core areas” and protecting undeveloped areas of forested vegetation to improve bird biodiversity in small tropical islands cities.

     

Developing a framework for stormwater management: leveraging ancillary benefits from urban greenspace

Managing stormwater and wastewater has been a priority for cities for millennia, but has become increasingly complicated as urban areas grow and develop. Since the mid-1800s, cites often relied on an integrated system of underground pipes, pumps, and other built infrastructure (termed gray infrastructure) to convey stormwater away from developed areas. Unfortunately, this gray infrastructure is aging and often exceeds its designed capacity. In an effort to alleviate issues related to excess stormwater, many urban areas across the United States are interested in using green infrastructure as a stopgap or supplement to inadequate gray infrastructure. Green infrastructure and other greenspace promote interception and/or infiltration of stormwater by using the natural hydrologic properties of soil and vegetation. Furthermore, there are numerous ancillary benefits, in addition to stormwater benefits, that make the use of greenspace desirable. Collectively, these ecosystem services can benefit multiple aspects of a community by providing benefits in a targeted manner. In this paper, we present a framework for balancing stormwater management against ancillary benefits of urban greenspace. The framework is structured around the Millennium Ecosystem Assessment ecosystem service categories: provisioning, cultural, regulatory, and supporting services. The purpose is to help communities better manage their systems by 1) allowing stakeholders to prioritize and address their needs and concerns within a community, and 2) maximize the ecosystem service benefits received from urban greenspace.

     

Urban biodiversity hotspots are not related to the structure of green spaces: a case study of tenebrionid beetles from Rome,Italy

At global and regional scales, area prioritisation is frequently done by the identification of hotspots based on species extinction risk. The logic of the hotspot identification has never been used in urban contexts. In this paper, the tenebrionid beetles (Coleoptera Tenebrionidae) of urban Rome were studied as an exercise to show how the hotspot approach can be profitably used in an urban area to identify priority areas for biodiversity conservation. For this, tenebrionid species from 16 green spaces were scored according to their vulnerability on the basis of their geographical distribution, habitat specificity and abundance. Species vulnerability scores were then used to calculate two indices of area prioritisation (the Biodiversity Conservation Concern and the Biodiversity Conservation Weight) for each green space. Values of these indices were correlated with site characteristics and compared with those obtained from other, more natural contexts. Except for distance to other sites, no significant correlation was found between conservation values and site characteristics, which indicates that the conservation importance of green spaces cannot be predicted on the basis of their geographical characteristics, but must be established on the basis of the species that they actually host. The importance of urban green spaces for biodiversity conservation may be questioned because of the large presence of ubiquitous and alien species in urban areas. Conservation values obtained for tenebrionids of green spaces in Rome are similar to those of various animal groups in more natural contexts and hence highlight the actual importance of green areas for insect biodiversity conservation.     

Mammal conservation in a changing world: can urban gardens play a role?

Urbanisation threatens biodiversity globally, yet some animal populations persist within urban landscapes. Conservation of urban wildlife has prioritised parks and remnant bushland as critical habitat and neglected the role that residential gardens offer for conservation. We explored the potential for residential gardens to assist the conservation of mammals using an online questionnaire administered to residents of two case study cities in Australia to identify how widespread mammals in cities can be, which garden features promote mammal presence, and if the features varied among species with different habitat requirements. From 649 responses we found that three mammal species with different habitat requirements occurred in residential gardens throughout the city landscape. Garden features promoting mammal presence were consistent with broad ecological and habitat requirements of each species, but differed among both species and regions. Our study demonstrates that residential gardens offer a valuable habitat for mammals, and that garden features could be manipulated to promote use of gardens by these species. By considering gardens in urban planning and management actions aimed at conserving urban wildlife, residential gardens offer additional habitat to parks, roadside vegetation and urban bushland, and can play a significant role in biodiversity conservation.

     

Renaissance of a rural artifact in a city with a million people: biodiversity responses to an agro-forestry restoration in a large urban traditional fruit orchard

The rural landscapes surrounding large cities are rapidly becoming incorporated into the urban environment. The most conspicuous changes involve green spaces, such as former agro-forestry systems like fruit orchards. In this paper, we assess the influence on biodiversity of restoring a large urban traditional fruit orchard as reflected by six selected taxa: plants, lichens, butterflies, beetles, orthopteroids and birds. The study was performed in Prague, which is the capital city of the Czech Republic and has more than a million inhabitants. We studied the effect of orchard renewal in 45 patches (15 for birds and 30 for other taxa). The majority of taxa responded positively to the restoration. The restoration had a significant positive effect on the species richness of lichens, butterflies and beetles. All taxa showed significantly altered species compositions, and the number of red-listed species increased. Orchards have a high potential for multi-functional use. Orchards are productive agro-forestry systems and host numerous possible human activities. Therefore, orchard restoration also has a social aspect. Moreover, our research in this artificial ecosystem revealed that its restoration increased the biodiversity and conservation potential of the associated areas.

     

Nested patterns in urban butterfly species assemblages: respective roles of plot management,park layout and landscape features

Increasing numbers of cities are currently developing sustainable policies aimed at promoting urban biodiversity and ecological dynamics through the planning of green networks and the implementation of more sustainable management practices. These human activities can strongly influence environmental factors on which the organization of ecological communities at different scales depends. Thus, it is of fundamental importance to understand the relative impact of local management, green space design and landscape features on the distribution and the abundance of species in urban areas. On the basis of 2 years of butterfly surveys in urban public parks within an extensive Mediterranean metropolitan area, Marseille (South-East France), the aim of this paper is to provide a better understanding of the effect of these three environmental scales (plot, park, landscape) on the composition and organization of species assemblages. Using variation partitioning and nestedness analysis on ecological data aggregated at plot-level and park-level respectively, we demonstrate the preponderant effect of landscape scale features on urban butterfly assemblages. Our results also highlight an important co-variation of plot management, park layout and urban landscape features, in their interaction with the community structure of urban butterflies. Although there is no significant species-area relationship, significantly nested patterns arise in species composition. Selective colonization appears as a driving force constraining the constitution of species assemblages within the city. However, a prospective study on adjacent more natural areas suggests that biotic limitations, interspecific competition and habitat filtering may play an important role if a larger portion of the urbanization gradient is explored, which remains to be investigated.     

Variability of bumblebee communities (Apidae,Bombini) in urban green areas

Based on current research the agglomerations are potentially desirable habitats for bumblebees. However, the relationship between the biodiversity of these bees and the green areas where they live is poorly understood. The aim of the study was to estimate the influence of green areas (ranging from 8 to 102 ha) of big cities on bumblebee species richness, composition, and the relative number of these insects. The studies were conducted within the administrative borders of the city of Wroc?aw (Poland) in 2011–2012 in 12 green areas such as parks, cemeteries and other places with trees and shrubs. Species richness and abundance of bumblebees was determined by direct observation during 30 min. The gathered materials were used to calculate how areas of urban green space affected qualitative and quantitative bumblebee community structure. In total, 13 species of bumblebees (Bombus Latr.) were recorded, of which 3 belonged to cuckoo bumblebees (Psithyrus subgenus). The share of the most similar groups was congregated in green areas not smaller than 30 ha. This was proved by analysis of qualitative structure (Sørensen index), quantitative structure (Renkonen index), and qualitative-quantitative structure (Cody’s index). The number of bumblebee species in the surveyed green areas (r = 0.7497) was decisive for the arrangement of the mutual similarity of group structure. Green urban areas should be created in a size of at least 30 ha. Such sites provide conditions for the most diversified bumblebee species communities. Sites smaller than 30 ha can play an important role as refuges, and allow migration to all pollinators.     

Are vascular epiphytes in urban green areas subject to the homogenization of biodiversity? A case study in the Brazilian Atlantic Forest

Urbanization is a disturbance process that can select species and result in biodiversity homogenization. Despite this, urban green areas shelter nature and are also important to human welfare. Epiphytes are an important functional group present in such areas, that are rarely studied. We evaluated the vascular epiphytic component in 26 urban green areas in the Brazilian Atlantic Forest and tested hypotheses related to the anthropogenic disturbances: 1) the community of epiphytes in urban green areas presents low richness and diversity of species; 2) there is low beta diversity due to flora homogenization represented by a reduced set of more tolerant species to disturbance. A total of 2288 trees (1563 representing phorophytes) and 110 epiphytic species were sampled. Six species were dominant, resulting in low diversity values, but some green areas had relatively high richness. The similarity found between the majority of the sampled areas suggests that epiphytic flora is subject to homogenization due to environmental filters. We found a high richness of species without adaptations to the epiphytic lifeform (accidental epiphytes) (42% of total sampled). Some results suggest that more comprehensive ecological and/or floristic studies about the epiphytes in the urban environment are necessary, such as Orchidaceae as the second richest family (since it is often poor in urban areas). More information about the species distribution patterns are necessary, both regarding the anthropized environments and the relationships with exotic or native phorophytes, as well as to enhance the knowledge of the ecological functions played by the epiphytes in these places.

     

Engaging urban nature: improving our understanding of public perceptions of the role of biodiversity in cities

In recognition of the value of biodiversity for cities and citizens, a number of international programs have been designed to help municipal governments sustain, protect, and augment the biodiversity and ecosystem services within their jurisdictions. A key component of these programs is public engagement, where citizens assume a more active role in maintaining urban biodiversity and associated ecosystem services. Yet, there are few studies which have as their focus public knowledge of the importance of nature, biodiversity, and ecosystem services in cities. To these ends, this study was conducted to develop a better understanding of how the public understands and interacts with urban biodiversity, particularly in comparison to subject matter experts. Using topics generated from expert interviews and the literature, an interview guide was developed for the general public, structured around the general themes of a definition of urban biodiversity, as well as the perceived benefits, costs, and threats related to urban biodiversity. While there were similarities in the responses of citizens and experts, some differences did emerge in terms of accounting for specific urban ecosystems, acceptable interventions to support and enhance biodiversity, and the character and extent of the cultural services derived from urban nature. Insights from this work can be used to inform education and information efforts for the public, as well as raise awareness among city planners and nature professionals of the array of urban ecosystem services recognized and made use of by the public.

     

Adaptive governance to promote ecosystem services in urban green spaces

Managing urban green space as part of an ongoing social-ecological transformation poses novel governance issues, particularly in post-industrial settings. Urban green spaces operate as small-scale nodes in larger networks of ecological reserves that provide and maintain key ecosystem services such as pollination, water retention and infiltration, and sustainable food production. In an urban mosaic, a myriad of social and ecological components factor into aggregating and managing land to maintain or increase the flow of ecosystem services associated with green spaces. Vacant lots (a form of urban green space) are being repurposed for multiple functions, such as habitat for biodiversity, including arthropods that provide pollination services to other green areas; to capture urban runoff that eases the burden on ageing wastewater systems and other civic infrastructure; and to reduce urban heat island effects. Because of the uncertainty and complexities of managing for ecosystem services in urban settings, we advocate for a governance approach that is adaptive and iterative in nature—adaptive governance—to address the ever changing social order underlying post-industrial cities and offer the rise of land banks as an example of governance innovation.     

Urban gradients alter the diversity,specific composition and guild distribution in tropical butterfly communities

The increasing expansion of the urban environment has been considered one of the causes of biodiversity loss. Due to the decrease of the quantity and quality of habitats, this process causes the restriction of organisms to sometimes-specific microhabitats. The patterns of variation of the biodiversity along the environmental gradients, however, are still poorly understood. This study evaluated how urban gradients affect the diversity, specific composition and frequency of guild of the community of butterflies. Butterflies were collected in three transects continuously traversing the urban, rural and preserved habitats using baited traps and by active sampling. We found that the urban gradient acts as an environmental filter in the butterfly communities, reducing the richness and frequency of guilds of frugivorous butterflies, structuring the community through turnover, but without effects in the abundance. This study shows the importance of the urban gradients approach in the butterfly community and its applicability in the management of urban areas in order to include native vegetation along the gradients, aiming at increasing environmental heterogeneity, to preserve butterflies and maintain the ecosystem services and network interactions in altered landscapes.

     

Woody plant species richness,composition and structure in urban sacred sites,Grahamstown, South Africa

Sacred sites are important not only for their traditional, spiritual or religious significance, but may also potentially be valuable for biodiversity conservation in human transformed landscapes. Yet, there has been little consideration of sacred sites in urban areas in this respect. Consequently, to better understand the ecosystem service and conservation value of urban sacred sites, inventories of their floral communities are needed. We examined the richness, composition and structure of the trees and shrubs in 35 urban churchyards and cemeteries in the City of Saints (Grahamstown). The combined area of urban sacred sites (38.7 ha) represented 2.2% of the city area and 13.6% of the public green space area. Species richness of woody plants was high, albeit dominated by non-native species. Levels of similarity among sites were low, indicating the effects of individual management regimens. There was no relationship between age of the site and measured attributes of the vegetation, nor were there any significant differences in vegetation among different religious denominations. However, the basal area and number of woody plants was significantly related to site size. These results indicate the significant heterogeneity of urban sacred sites as green spaces within the urban matrix. The significance of this heterogeneity in providing ecosystem services to users of sacred sites and the broader urban communities requires further investigation.