URban Biotopes of Aotearoa New Zealand (URBANZ) (I): composition and diversity of temperate urban lawns in Christchurch

Christchurch urban lawns are dominated by non-native grasses and forbs. However, we document considerable plant diversity; the total number of species encountered in our 327 sampled lawns was 127, although 80 species occurred in <2% of lawns. Seven distinct lawn communities were identified by Two-Way INdicator SPecies ANalysis using occurrence of 47 species that occurred in > 2% of lawns. Our ability to explain variation in species composition was surprisingly good and indicates intensity of lawn maintenance such as frequency of mowing, irrigation, fertiliser, and herbicide use and whether clippings are removed or not plays the major role. Species richness significantly declines with an increase in total area of contiguous lawn, leaf litter cover, the presence of grass clippings, and on loamy soil. Hence, park lawns with coarser management had lower species richness than residential lawns. Native species were more prevalent in well tended residential lawns, where more frequent mowing and removal of clippings or litter build-up diminishes shoot competition or shading. There is tremendous potential for more native species in New Zealand lawns which would contribute substantially to the conservation of endangered lowland herbaceous flora.     

Floristic composition and richness of urban domestic gardens in three urban socioeconomic stratifications in the city Heredia,Costa Rica

The urban domestic gardens that are part of urban green areas have become a very important element due to their ecosystematic benefits and the poor information about them. Information regarding the floristic richness of three socioeconomic stratifications from the city of Heredia, Costa Rica is presented in this study. Research was conducted from October 2011 to May 2012. Information was collected using a semi-structured questionnaire, which was provided to 61 garden owners, including the owner’s personal data, time living in the dwelling, area of the property and the garden, variation of its size in time and the garden’s location on the property. In addition, a floristic inventory was conducted in each garden as well as a taxonomic identification of all plants. Vegetative forms and substrates were also noted. Characteristics of the gardens and their owners were then analyzed to determine the relationship of species richness between the three sites studied. The number of plant species recorded was 618, corresponding to 102 families (19 % exotic). Regarding geographical origin, 64, 4 % were exotic, 35,6 % native. Orchidaceae (93 spp.) and Araceae (39 spp.) were the most dominant families. The variables of garden area and location and owner's age determined the garden’s floristic richness. In addition, the garden’s area and vegetative forms varied among the sites studied probably due to socioeconomic differences among them. Diversity of pioneer plants increased with the garden’s area. In sum, this research provides important floristic information of domestic gardens, since, being private property, data about them is scarce and municipal management guidelines are nonexistent.     

Vulnerability of native and exotic urban birds to housing densification and changing gardening and landscaping trends

Variation in densities of native and exotic urban-adapted birds across suburban habitats may reveal levels of adaptation that predict vulnerability to habitat modification over time. We investigated the densities of the eight most common exotic and native birds in a southern hemisphere city across suburbs of varying housing density, vegetative cover and garden vegetation characteristics to determine whether avian population densities are maintained across variations in habitat structure that mirror current temporal trends in land use and landscaping/gardening preferences. We also tested whether densities of birds in residential areas bordering undeveloped patches were higher due to spill-over from patches. Densities of all four native species were highest in undeveloped patches and these species were largely absent from high density housing areas. While densities of two exotic species (blackbird, song thrush) were slightly higher in suburbs with greater vegetation complexity, they were still present in higher density housing suburbs in significant numbers. Common starlings were similarly abundant across all residential habitats and house sparrow density increased in higher density housing areas. Blackbirds appeared least vulnerable to housing densification and landscaping and gardening trends because they were abundant across all habitats, including undeveloped patches. Densities of native birds were higher in residential areas bordering undeveloped patches indicating spill-over was occurring and emphasising the role that patches could play in boosting city-wide native bird abundances. Native birds are vulnerable to reductions in garden size and vegetation complexity that should have little or positive effects on populations of common exotic species.     

Baseline biodiversity surveys of the soil macrofauna of London’s green spaces

A serious barrier to our understanding of urban ecosystems is a lack of information on the ecology of soils organisms of green spaces within large cities. This study addresses this gap by providing baseline survey data on the biodiversity of soil macrofauna in urban parks and domestic gardens of London, UK. In April and June 2004, the soil macrofauna were handsorted from soil cores in eleven parks and gardens of various sizes in central London. Five taxa were identified to species (Lumbricidae, Isopoda, Diplopoda, Chilopoda and Formicidae). The biodiversity value of the two main habitats (horticultural borders and mown grass lawns) was assessed and the influence of a range of environmental factors on species density (number of species per unit area) examined. The species densities of the studied soil invertebrates in the urban gardens were comparable with those found in natural ecosystems, although plant borders contained significantly more species than lawns. Borders had higher levels of plant nutrients, higher floristic diversity and lower levels of micronutrients and heavy metals than lawns. Significant predictor variables of species densities in the plant borders were the percentage of leaf litter cover, sampling month and soil pH. Species densities in the lawns were significantly correlated with the distance of the samples from the edge of the lawn.     

Changes through time in soil Collembola communities exposed to urbanization

The study aimed to assess if long-term exposure to urbanization changes the structure and composition of soil collembolan communities in urban green components (street lawns and park lawns) and in all urban green. Species diversity metrics, rarefaction, species richness estimators (Chao 1 and ACE) and multivariate analysis were used for the comparison of changes in community structure and diversity pattern over ca 30 years’ time span. Our results clearly demonstrate a shift, through time, in Collembola community composition and structure in an urban ecosystem and confirm that there is a linkage between long-term exposure to urbanization and changes in collembolan communities. Long-term urbanization led to erosion in species diversity and the formation of species-poor communities, species replacement, loss of specialized forms and promoted the invasion of exotic species. However, we show that the time span considered produced significant differences in diversity attribute values for the collembolan communities from street lawns and insignificant differences in park lawns, also we noted lack of significant differences in collembolan abundance across the two urban green components. The observed temporal changes in collembolan communities indicate that their response to disturbances in urban settings and selecting species is shaped by multiple processes. We conclude that more resistant collembolan communities were found over time in less stressed urban greening components such as park lawn soils compared to street lawn soils.     

Influence of Argentine and coastal brown ant (Hymenoptera: Formicidae) invasions on ant communities in Perth gardens,Western Australia

A survey examined the influence of Argentine (Linepithema humile (Mayr)) and coastal brown ant (Pheidole megacephala (Fabricius)) populations on other ants in the Perth metropolitan area, Western Australia. Twelve gardens (yards) were sampled; four infested by Argentine ants, three infested by coastal brown ants, and five controls where these two tramp ants were absent or collected only as isolated strays. Collection methods used were pitfall trapping and hand collection. A total of 27 species was recorded. Eight of these species, including the above two tramp ants, are non-native to the region. Pheidole megacephala was the most abundant species, comprising 95.4% of ants in P. megacephala–infested gardens. Linepithema humile comprised 92.1% of ants in L. humile–infested gardens. Other common ants were the native Iridomyrmex chasei and the introduced species Tetramorium simillimum and Tetramorium bicarinatum. The cryptic introduced species Cardiocondyla nuda and Tetramorium simillimum appear able to persist in small numbers in L. humile–infested gardens, but virtually no other ant species occurred where the coastal brown ant was well established. Mean richness, diversity, and evenness were significantly different between P. megacephala–infested and control gardens, and significantly different between L. humile–infested and control gardens. Diversity and evenness, but not richness, were significantly different between P. megacephala–infested and L. humile–infested gardens. Ordination analysis revealed that the three types of gardens had very different ant community profiles.     

Diversity,abundance, and species composition of ants in urban green spaces

Urbanization threatens biodiversity, yet the number and scope of studies on urban arthropod biodiversity are relatively limited. We sampled ant communities in three urban habitats (forest remnants, community gardens, vacant lots) in Detroit and Toledo, USA, to compare species richness, abundance, and species composition. We measured 24 site characteristics to examine relationships between richness and composition and habitat patch size, vegetation, and urban features. Ant richness was higher in forests (26) than in gardens (14) and intermediate in vacant lots (20). Ant richness in gardens and vacant lots negatively correlated with abundance of an exotic ant species (Tetramorium caespitum); thus this ant may affect native ant richness in urban habitats. Ant composition differed with habitat type, and abundance was lowest in forests. Site characteristics varied with habitat type: forests were larger, had more woody plants, higher woody plant richness, more branches, and leaf litter whereas lots and gardens had more concrete and buildings. Vacant lots had taller herbaceous vegetation, and gardens had higher forb richness, density, and more bare ground. Differences in vegetation did not correlate with ant richness, but several vegetation factors (e.g. patch size, number and size of trees, leaf litter, and amount of concrete and buildings) correlated with differences in ant species composition. Additional factors relating to soil, nests, or microclimatic factors may also be important for urban ant communities. Implications for biodiversity conservation in urban ecosystems are discussed.     

Urban areas as hot-spots for introduced and shelters for native isopod species

Isopod assemblages were studied in Budapest, capital of Hungary. The analyses of literary and field data revealed a high species richness (28 species), compared to the total species number (57) in Hungary. Habitats characteristics for the city were categorized as native forests, urban forests, gardens of Buda, gardens of Pest, public parks, densely built-up areas and botanical gardens. We hypothesized that isolated and diverse habitat patches in the city matrix of Budapest support the introduction and establishment of exotic species and the survival of native ones. The composition of assemblages varied among sampling sites, but were characteristic for the biotope categories. We concluded that forests, parks and gardens play an important role in the survival of native isopod populations. Species numbers were highest in the gardens of Buda and in the botanical gardens (both 17 species). The overall presence of cosmopolitan and disturbance-tolerant species indicates an ongoing homogenization process.     

Estimating urban lawn cover in space and time: Case studies in three Swedish cities

Lawns are considered monocultures and lesser contributors to sustainability than diverse nature but are still a dominating green area feature and an important cultural phenomenon in cities. Lawns have esthetical values, provide playground, are potential habitat for species, contribute to carbon sequestration and water infiltration, but also increase pesticides, fertilization, are monocultures and costly to manage at the same time. To evaluate the potential impact of lawns, whether positive or negative, it is of interest to estimate the total lawn cover in cities and its change over time. This is not a straightforward process, e.g., because many lawns are small and covered by trees. In this study we review the existing literature of lawn cover in cities and the different methodologies used for cover estimation. We found both pros and cons with NDVI and LiDAR data as well as manually interpreted aerial photos. The total cover of lawns in three case study cities was estimated to 22.5%. By extrapolating these percentages to all Swedish cities lawn cover was estimated to 2589 km² (0.6% of the terrestrial surface). The approximated total municipal management cost of lawns in all Swedish cities was 910,000,000 USD/ year. During 50 years lawn area almost doubled in relative cover and 56% of them were continuously managed. Since lawns constitute large parts of the urban greenery and are costly to manage it is highly relevant to consider their social, ecological and cultural value compared to alternatives, e.g., meadows with less intensive management.     

City dwelling wild bees: how communal gardens promote species richness

Urban areas consist of wide expanses of impervious surfaces which are known to negatively affect insect biodiversity in general, but green spaces within cities have the potential to provide necessary habitat and foraging resources. Although, communal gardens were primarily intended to provide fresh, regional food to denizens, these green islands also host a surprisingly high number of wild bee species.

The gardens were characterized based on structural elements such as flower frequency, the relative percentage of lawn, trees, shrubs, planted crops and infrastructure (e.g. seating possibilities or garden houses). Further, the effects of different landscape structures surrounding the gardens and distance to the city center were analyzed on the total wild bee species richness and functional traits. Focusing on these putative influencing factors, statistical analyses calculating random decision forests along with generalized linear mixed models were applied. With 113 observed wild bee species, communal gardens provide habitat for a quarter of all known species in Vienna. In conclusion, results revealed that only elements within the gardens had an effect on species richness, with flower frequency as the major positive driver. The examined communal gardens promote and conserve wild bees independent from the location within the city or garden size. Furthermore, these green patches are important sanctuaries, hosting rare and threatened species as well as remarkably special wild bee communities.

     

Contrasting soil nitrogen dynamics across a montane meadow and urban lawn in a semi-arid watershed

Urbanization substantially increases nitrogen (N) inputs and hydrologic losses relative to wildland ecosystems, although the fate of N additions to lawns and remnant grasslands remains contested. In montane semi-arid ecosystems, N cycling is often closely coupled to snowmelt (the dominant period of infiltration) and snow cover, which impact soil temperature and moisture. Here, we compared soil N dynamics between a fertilized and irrigated urban lawn and nearby riparian meadow in Salt Lake City, Utah during a snow manipulation experiment. Snow removal increased freeze/thaw events but did not affect N pools, microbial biomass, denitrification potential, or soil oxygen (O₂). Mineral N was similar between sites despite lawn fertilization, but dissolved organic N (DON) was four-fold greater (2.1 ± 0.1 mg N l^?1) in lawn soil water. Infiltration was lower in the lawn subsoil, and leaching losses (modeled with Hydrus) were small at both sites (< 2 kg N ha^?1 y^?1) despite substantial lawn fertilization. Lawn soil O₂ fluctuated between 20.9 and 1.6 % following snowmelt and irrigation, but remained near 20 % in the meadow; the lawn had more reducing microsites as indicated by iron speciation. Post-snowmelt potential denitrification was six-fold greater in the lawn than the meadow. Lawns can potentially provide hotspots of denitrification in a semi-arid landscape that exceed some natural riparian ecosystems, whereas DON may represent an increasingly important form of N loss from lawns.     

Soil-atmosphere exchange of carbon dioxide,methane and nitrous oxide in urban garden systems: impact of irrigation,fertiliser and mulch

Urban green spaces provide important ecosystem services, such as amenity, biodiversity, productivity, climate amelioration, hydrological and biogeochemical cycling. Intensively managed urban gardens can sequester carbon through vegetation growth and soil C increase, but may experience nitrous oxide (N₂O) emissions and reduced soil methane (CH₄) uptake from irrigation and fertiliser use. Soil atmosphere exchange of N₂O, CH₄ and carbon dioxide (CO₂) was measured in lawn and wood chip mulched garden areas in Melbourne, Australia in winter, spring and summer under various water and fertiliser regimes. Gas exchange before and after lawn fertiliser application was measured continuously for three weeks using an automated chamber system. Applying fertiliser led to a peak N₂O emission of >60 μg N m⁻² h⁻¹, but overall only weekly irrigation (10 mm) significantly increased mean soil N₂O emissions above that in other treatments. Under mulch, mean soil N₂O emissions (14.0 μg N m⁻² h⁻¹) were significantly smaller than from irrigated lawn (27.9 μg N m⁻² h⁻¹), whereas mean soil CH₄ uptake under mulch (−30.7 μg C m⁻² h⁻¹) was significantly greater (p < 0.01) than in any lawn treatment. Lawns were either a weak CH₄ sink or source. Soil C density (0–25 cm) under mulch (12.5 kg C m⁻²) was greater that under lawn (8.0 kg C m⁻²). On a carbon dioxide equivalent (CO₂-e) basis, soil N₂O emissions offset the benefits of soil CH₄ uptake. Mulched garden areas provide greatest C sequestration potential in soil and vegetation and the smallest non-CO₂ emissions, as soil CH₄ uptake offsets a large fraction of soil N₂O emissions. Results of this study suggest that reducing the irrigation and fertiliser application to lawns can help mitigate GHG emissions from urban garden systems, and increasing the area of mulched perennial garden beds can also provide net GHG benefits; however, this needs to be tested in other garden systems with different soil types and environmental conditions.     

Homestead greening is widespread amongst the urban poor in three medium-sized South African towns

Trees in urban areas are important sources of ecosystem services and benefits. In most towns the bulk of urban biodiversity, and trees specifically, are found in homestead gardens. But there is only limited understanding of the tree holdings in such gardens, and how they vary within and between towns, especially for developing countries where rapid urbanisation and high poverty influence the use of and reliance on land and local resources. We report on the nature of tree holdings in private gardens of poorer suburbs in three medium-sized towns along a gradient of decreasing mean annual rainfall in northern South Africa. A total of 3 217 trees were enumerated across 450 randomly selected homesteads. Most (90 %) households had at least one tree on their homestead, with an average of 7.7?±?6.1 trees. Most householders had planted the trees themselves. The density of trees declined along the moisture gradient. Within towns, tree density was positively related to garden size, which in turn was related to relative affluence and age of the suburb. Newer and poorer suburbs had the fewest trees per household. Sixty-two tree species were recorded, which were dominated by alien species, especially fruit trees. There was no relationship between the moisture gradient and tree species richness per household, but within towns there was a difference between suburbs, being lowest in the newest suburbs. Numbers of trees and species per household was positively related to age of the household head.     

Insects on urban plants: contrasting the flower head feeding assemblages on native and exotic hosts

Exotic plant species very often comprise a large proportion of urban floras. Because herbivorous insects depend on the presence of suitable host plants to maintain their populations, it is imperative to elucidate the relative importance of native and exotic hosts to understand the response of herbivorous guilds to urbanization. By using a plant-herbivore system composed of Asteraceae hosts and flower-head endophagous insects, we investigated whether the diversity and composition of herbivorous insects differs between native and exotic host-plant species in an urban environment. Although we found only seven exotic Asteraceae among the 30 species recorded, the overall abundance of exotics was considerably greater than that of native host plants. Overall, the exotic host species supported a small subset of the herbivore assemblage found on the native ones. The number of herbivore species per host species was significantly higher among the native plants, but we did not find a difference in herbivore abundance. Moreover, the higher taxonomic composition of herbivores on exotic Asteraceae was reduced, being composed of only three genera and two families from a total of 16 genera and six families of herbivores. These results provide support for the idea that plants outside of their original geographic distribution have lower loads of enemies than phylogenetically related native species. Our findings indicate that native host plants in urban areas play a critical role in supporting the native herbivorous insect fauna.     

Implementing integrated pest management in professional lawn care: a case study

Human choices regarding land cover management practices may influence ecosystem services provided by urban green spaces. We conducted a 2-year study to compare biological (weed, insect, and disease), aesthetic (lawn quality), and economic (lawn care program cost) attributes of an integrated pest management (IPM) program, in which pesticides are applied on the basis of treatment thresholds, with a standard program, in which pesticides are applied on a calendar basis without pest monitoring. Both programs were managed by a professional lawn care operator. Although weed incidence was low, the IPM program had significantly more lawns with weed presence than the standard program during 2005 and 2006. However, only 21% of the IPM lawns required herbicide applications in 2005, and none exceeded the treatment threshold (5% weed cover) in 2006 as compared to 100% of the standard program lawns being treated for weeds in both years. The IPM program also had significantly more lawns with insect damage than the standard program during June 2005 and August 2005, but not September 2005 and throughout 2006. Only 28% of the IPM lawns required insecticide applications in 2005 and none exceeded the threshold (5% insect damage) in 2006 whereas all of the lawns in the standard program received insecticide treatments in both years. Rhizoctonia blight was present on some of the lawns, but was not a common problem. Although lawn quality was high for both programs (>8, on a scale of 1–9), it was significantly higher for standard than for IPM program lawns during 2005, and June 2006 and September 2006, but not August 2006. The annual lawn management costs were lower for the IPM (281.50) than the standard program (281.50) than the standard program (458.06). Thirty one percent of the IPM program customers who continued with the study in 2006 did so because they were satisfied with the IPM program. Among those who did not continue with the program, 33% cited weed or insect problems, while 33% expected better results. The implications of these findings for implementation of IPM in professional lawn care are further discussed.     

Vegetation on and around large-scale buildings positively influences native tropical bird abundance and bird species richness

High population growth in the tropics is driving urbanisation, removing diverse natural ecosystems. This is causing native species to suffer while introduced synanthropes flourish. City planners are developing urban greenspace networks, in part trying to address this issue. Architects contribute to these greenspace networks by designing elevated and ground level green spaces on large-scale buildings. However, little evidence is available on whether building green spaces support native fauna. This is true for birds in tropical Singapore that support important ecosystem services and have existence value. Therefore, in this study, we conducted bird surveys and statistical analyses to determine, if and how vegetation on three building green space types (ground gardens, roof gardens and green walls) have a positive impact on native or introduced bird species. We found that elevated greenery (roof gardens and green walls) on large-scale buildings supported a higher richness of birds and abundance of urban native birds than control roofs and walls without vegetation. Ground gardens supported similar levels of native species as roof gardens but also a larger proportion of generalist synanthropes. However, we found no tropical forest habitat specialists across any space type. Therefore, we recommend roof gardens and ground gardens as a potential space for urban natives outside of a less competitive ground-level urban environment. Our study also found certain building design elements (height of elevated space, presence of specific plants) supported different species groups. Therefore, we suggest that these ecological requirements for different species groups are considered when designing a building’s green space.

     

Bee-friendly community gardens: Impact of environmental variables on the richness and abundance of exotic and native bees

With their abundant floral resources, urban community gardens have the potential to play an important role in pollinator conservation. At the same time, the gardens themselves are dependent upon the pollination services provided by insects. Thus, understanding the variables that can increase bee richness or abundance in community gardens can contribute to both urban agriculture and pollinator conservation. Here we examine the impact of several environmental variables on bee abundance and diversity in urban community gardens in Sydney, Australia. We used hand netting and trap nests to sample bees in 27 community gardens ranging from inner city gardens with limited surrounding green space, to suburban gardens located next to national parks. We did not find strong support for an impact of any of our variables on bee species richness, abundance or diversity. We found high abundance of a recently introduced non-native bee: the African carder bee, Afranthidium repetitum (Schulz 1906). The abundance of African carder bees was negatively correlated with the amount of surrounding green space and positively correlated with native bee abundance/species richness. Our results highlight the seemingly rapid increase in African carder bee populations in inner city Sydney, and we call for more research into this bee’s potential environmental impacts. Our results also suggest that hard-to-change environmental factors such as garden size and distance to remnant forests may not have a strong influence on native bee diversity and abundance in highly urbanized area.     

Ecology of urban lawns under three common management programs

Turfgrass lawns are a central part of urban and suburban landscapes throughout North America and are often managed using repeated applications of chemical fertilizers and pesticides. These inputs are expensive and may negatively affect ecological processes in lawns. Therefore, we evaluated the influence of three most common lawn care programs on ecological characteristics of turfgrass lawns. Twenty-eight home lawns, separated into 3 groups based on the lawn care program (professional, do-it-yourself [DIY], and no-input), were studied. Data on lawn quality, weed and insect infestation, disease incidence, soil nematode community, soil nitrogen pools, microbial biomass (MBN), and soil organic matter (SOM) were collected. Results indicated that professional lawn care resulted in the highest aesthetic lawn quality mainly due to better weed control, compared to DIY and no-input programs. However, professional and DIY programs negatively affected MBN and SOM pools and enhanced disease (rust) severity. No significant differences in soil nematode population and nematode community indices across the three programs were found, indicating no differences in net ecosystem productivity among the three programs. Overall, soil nematode food web in turfgrass lawns represented a disturbed food web compared to natural grasslands and forest ecosystems, irrespective of the lawn care program used.     

Ecosystem services provided by urban spontaneous vegetation

Spontaneous vegetation colonizes large areas in and around cities. These unmanaged areas are considered to have low economic value or indicate dereliction, but recent research suggests that these can contribute valuable ecosystem services. This study evaluates indicators of ecosystem services in three habitats: urban spontaneous vegetation (USV), managed lawns, and semi-natural urban forest, in Halifax, Nova Scotia. USV had higher indicator values for habitat provisioning (plant species diversity, invertebrate abundance and taxonomic diversity) than the other habitats. Indicators of climatic regulatory services (albedo and leaf area index) in USV were similar to those in lawn habitats. Organic carbon content of the soils, an indicator of carbon storage, was lowest in USV but only marginally lower than in lawns. Standing biomass, an indicator of production services, was lowest in USV but lawn production may have been overestimated. While USV sites are usually transitory components of the urban landscape, they deserve further consideration due to their provision of ecosystem services, in some cases to a greater extent than conventionally valued urban habitats.     

Impacts of urbanisation on the native avifauna of Perth, Western Australia

Urban development either eliminates, or severely fragments, native vegetation, and therefore alters the distribution and abundance of species that depend on it for habitat. We assessed the impact of urban development on bird communities at 121 sites in and around Perth, Western Australia. Based on data from community surveys, at least 83 % of 65 landbirds were found to be dependent, in some way, on the presence of native vegetation. For three groups of species defined by specific patterns of habitat use (bushland birds), there were sufficient data to show that species occurrences declined as the landscape changed from variegated to fragmented to relictual, according to the percentage of vegetation cover remaining. For three other groups (urban birds) species occurrences were either unrelated to the amount of vegetation cover, or increased as vegetation cover declined. In order to maximise the chances of retaining avian diversity when planning for broad-scale changes in land-use (i.e. clearing native vegetation for housing or industrial development), land planners should aim for a mosaic of variegated urban landscapes (>60 % vegetation retention) set amongst the fragmented and relictual urban landscapes (<60 % vegetation retention) that are characteristic of most cities and their suburbs. Management actions for conserving remnant biota within fragmented urban landscapes should concentrate on maintaining the integrity and quality of remnant native vegetation, and aim at building awareness among the general public of the conservation value of remnant native vegetation.