Patterns of human disturbance and response by small mammals and birds in chaparral near urban development

We report on the extent of disturbance (including habitat alteration and road and trail proliferation) in chaparral near urban development and analyze the effects of disturbance on small mammal and resident bird species. Disturbance patterns were evaluated in a 6700 ha study area in southern California: effects on mammals and birds were investigated by analyzing relationships between vegetation structure and animal species richness and abundance. Disturbance was prevalent throughout the study area and included extensive human-altered habitat (from past human activities such as vegetation clearing, human-caused fires, refuse dumping, and vegetation trampling) and 157 km of roads and trails. A nonsignificant trend was found between human-altered habitat and proximity to development, but human-altered habitat was significantly associated with roadway proximity. Trails were also more frequent near urban development and roads. Small mammals responded strongly to disturbance-related vegetation changes, while birds showed little or no response. Mammals endemic to chaparral vegetation were less diverse and abundant in disturbed sites, whereas disturbance-associated species increased in abundance. Close proximity of urban development to natural areas resulted in alteration of natural habitat and proliferation of roads and trails. Variation in life history traits between birds and mammals may affect response to disturbance and influence persistence if disturbance continues. Conservation efforts must recognize the potential for habitat damage and associated declines in native animal species caused by disturbance near urbanization and implement strategies to reduce these threats.     

Bird community responses to different urban conditions in La Paz,Bolivia

Urbanization alters ecosystems worldwide, but little is known about its effects in the Neotropical region. In the present research we examined the relative influence of different levels of urbanization and of some urban development measures on bird species richness, abundance and composition. We surveyed 104 observation stations at which we collected data on the relative abundance of bird species, and also data on seven environmental variables as measures of urban development and human activity. We registered 57 native bird species. Bird species richness and bird abundance increased with lower urbanization levels. Both variables were positively related to vegetation cover and native vegetation, and negatively to built-up cover, abundance of Rock Pigeon (Columa livia), pedestrian rate and car rate. A canonical correspondence analysis produced a significant model that explained 37% of the total variation in species data. This analysis segregated bird species along two important gradients: urbanization and elevation. The most urbanized areas were dominated by a few synanthropic species tolerant to human disturbance, such as Rufous-Collared Sparrow (Zonotrichia capensis), Eared Dove (Zenaida auriculata) or Chiguanco Thrush (Turdus chiguanco). Areas with lower urbanization levels had more species typically associated with native vegetation, such as Plain-Mantled Tit-Spinetail (Leptasthenura aegitaloides) or D’Orbigny’s Chat-Tyrant (Ochthoeca oenanthoides), among others. Elevation had a significant influence in structuring bird communities, with some species restricted to higher elevations and some to lower elevations. Although changes in elevation had an important influence, urbanization had a stronger effect on structuring bird communities. This study provides valuable information and an important baseline for future studies.     

Birds biodiversity in urban and periurban forests: environmental determinants at local and landscape scales

A significant decline in biodiversity is associated with the current and upcoming degree of urbanization. A challenging strategy to address this conflict is to make urban growth compatible with biodiversity protection and in this context urban parks can play a crucial role. Urban systems are highly dynamic and complex human-shaped ecosystems, where the relationship between species and environment may be altered and make the preservation of biodiversity within them a challenging goal. In this study, we analysed how different environmental features affect bird biodiversity in one of the most urbanized areas of Italy (the metropolitan area of Milan) at different spatial scales. Bird surveys were conducted in fifteen urban and peri-urban parks and environmental variables at landscape and local scale recorded. Results showed that a mixture of land covers and the presence of water bodies inside urban parks favoured species occurrence and abundance at landscape scale, but a surrounding dense urban matrix deflated biodiversity. At local scale, woodland cover and presence of water bodies were key determinants in ensuring overall high biodiversity but local-specific vegetation management produced an unusual pattern for forests species. In particular, the maintenance of large trees may not result in biodiversity support for forest bird species if large trees are not located in woodland areas with a significant tree density. To understand biodiversity patterns and provide useful information for urban planning and design, we need to provide insights into species/environment relationships at multiple scales in the urban environment.     

Nest design in a changing world: Great tit <Emphasis Type="Italic">Parus major</Emphasis> nests from a Mediterranean city environment as a case study

Investigations of urbanization effects on birds have focused mainly on breeding traits expressed after the nest-building stage (e.g. first-egg date, clutch size, breeding success, and offspring characteristics). Urban studies largely ignored how and why the aspects of nest building might be associated with the degree of urbanization. As urban environments are expected to present novel environmental changes relative to rural environments, it is important to evaluate how nest-building behavior is impacted by vegetation modifications associated with urbanization. To examine nest design in a Mediterranean city environment, we allowed urban great tits (Parus major) to breed in nest boxes in areas that differed in local vegetation cover. We found that different measures of nest size or mass were not associated with vegetation cover. In particular, nests located adjacent to streets with lower vegetation cover were not smaller or lighter than nests in parks with higher vegetation cover. Nests adjacent to streets contained more pine needles than nests in parks. In addition, in nests adjacent to streets, nests from boxes attached to pine trees contained more pine needles than nests from boxes attached to other trees. We suggest that urban-related alterations in vegetation cover do not directly impose physical limits on nest size in species that are opportunistic in the selection of nesting material. However, nest composition as reflected in the use of pine needles was clearly affected by habitat type and the planted tree species present, which implies that rapid habitat change impacts nest composition. We do not exclude that urbanization might impact other aspects of nest building behaviour not covered in our study (e.g. costs of searching for nest material), and that the strengths of the associations between urbanization and nest structures might differ among study populations or species.     

Forest bird communities across a gradient of urban development

This study examined native bird communities in forest patches across a gradient of urbanization. We used field data and multivariate statistical techniques to examine the effects of landscape context, roads, traffic noise, and vegetation characteristics on bird community composition in the North Carolina Piedmont (U.S.A.). Landscape-level variables, particularly those related to urbanization, were most important in structuring forest bird communities. Specifically, we found that road density and amount of urban land cover were the best predictors of species composition. We found that urban and rural bird communities were quite distinct from each other. Rural communities had more long-distance migrants and forest interior species but species richness did not differ between the communities. Our results suggest some specific guidelines to target bird species of interest both inside and outside of urban areas. For example, if increasing numbers of migratory species is of primary concern, then conservation areas should be located outside of urban boundaries or in areas with low road density. However, if maximizing species richness is the focus, location of the conservation area may not be as important if the conservation area is surrounded by at least 50 m of forest habitat in all directions.     

Are plant communities mainly determined by anthropogenic land cover along urban riparian corridors?

Often used as a mitigation tool to landscape fragmentation, urban riparian corridors also suffered from the effects of the urban expansion. This study explored the relationships between plant riparian communities and two major environmental variables (land cover, soil characteristics) and analyzed the floristic change along an urbanization gradient. Fifteen sites were surveyed on both riverbanks of two riparian corridors characterized by contrasting water regimes in Strasbourg, North Eastern France. Data of spontaneous species abundance was collected from 180 quadrats using (i) all plant species, (ii) herbaceous stratum and (iii) ligneous stratum (bush and tree). The diversity and compositional patterns of riparian plant species were analyzed within each corridor according to three levels of urbanization (urban, suburban, peri-urban). Relationships between riparian communities, land cover and soil chemical properties (pH, nitrogen and carbon content, moisture) were established by between-class co-inertia analyses. Land cover emerged as the main factor explaining changes in riparian communities along the rural–urban gradient while soil chemical properties discriminate water stress and fluvial dynamics of the two corridors. Similar compositional patterns were found within the most urbanized sites with the establishment of ubiquitous species. The herbaceous stratum is best linked to the level of urbanization, whereas the tree stratum is primarily correlated with corridor attributes (hydrological regime, soil properties). Although riparian species and communities are mainly determined by land cover, urban riparian corridors maintain native biodiversity up to the urban center.     

The influence of small urban parks characteristics on bird diversity: A case study of Petaling Jaya,Malaysia

The capacity of small urban park to serve as urban habitats are rarely explored. This study analyses the characteristics of small urban parks and their potential to support urban biodiversity and ecological functions. Nine small urban parks were studied in Malaysia in August and September 2014 using the combined field survey method of structured observation and field measurements. The measured variables were divided into three broad categories of physical characteristics, species richness and human factors. Bird species richness and abundance were used as the indicators for assessing biodiversity. Pearson correlations and multiple regressions were conducted to analyse the relationships between variables and to identify which variables had a significant effect on bird species richness and abundance. The results demonstrated that park area and vegetation variables ( e.g. the percentage of tree canopy cover, open grass/ground, native-exotic plants) are the important predictors of bird species richness and abundance. The percentage of canopy covers (negative relation) and park area (positive relation) are the best predictors of bird species richness in small urban parks. Meanwhile, the best predictors for bird abundance are the percentage of canopy covers (negative relation) and native vegetation species (positive relation). Human activities and park surroundings have a marginal effect on the presence of bird species in small parks. Based on the findings, we provide two general recommendations that could probably increase bird diversity in small urban parks: (1) the park development and management plan should incorporate a social-ecological approach that can benefit both city-dwellers and bird species, and (2) findings from the study should be used to rethink the planting design and composition of especially newly established small urban parks.     

Impact of urban structure on avian diversity along the Truckee River, USA

Urban riparian habitats are potentially important resources for native birds in arid ecosystems. Most studies have assessed the value of urban riparian habitat in terms of vegetation and natural resources; however, the surrounding land use and infrastructure may determine the viability of urban habitat. We studied the impact of urban structure, the combination of land use, infrastructure and vegetation variables that work together to shape the urban environment, on avian riparian habitat in the Truckee Meadows, Nevada, USA. Land use and infrastructure explained avian species richness and abundance better than local vegetation alone, but community resemblance was more strongly correlated to vegetation. Avian species guilds responded differentially to surrounding land use, suggesting there may be a functional difference between land use types. The best models for bird diversity used urban structure (both land use and vegetation) to describe potential habitat. Urban structure describes urban habitat in ways that vegetation variables alone cannot. Studies that ignore land use and infrastructure and other socioeconomic variables are likely missing key functional differences within urban ecosystems, and may miss the potential for compatible development that encourages both biodiversity and urban growth.     

Defining urban,suburban, and rural: a method to link perceptual definitions with geospatial measures of urbanization in central and eastern Massachusetts

Developing greater understandings of socio-ecological relationships across urbanizing areas is increasingly recognized as important for the conservation and management of natural resources in a variety of development contexts. Efforts to do so have been hindered by a lack of consistent measures of urbanization and the challenge of integrating socio-cultural characteristics into definitions of urban. We present a novel method for linking perceptual definitions of urban, suburban, and rural to geospatial characteristics and demonstrate how the method can be used to map urban, suburban, and rural areas at multiple scales in central and eastern Massachusetts. Our method can facilitate comparative approaches to urban ecology, be used to scale up socio-ecological studies, and inform conservation research and practice in urbanizing areas.     

Diversity,abundance, and species composition of ants in urban green spaces

Urbanization threatens biodiversity, yet the number and scope of studies on urban arthropod biodiversity are relatively limited. We sampled ant communities in three urban habitats (forest remnants, community gardens, vacant lots) in Detroit and Toledo, USA, to compare species richness, abundance, and species composition. We measured 24 site characteristics to examine relationships between richness and composition and habitat patch size, vegetation, and urban features. Ant richness was higher in forests (26) than in gardens (14) and intermediate in vacant lots (20). Ant richness in gardens and vacant lots negatively correlated with abundance of an exotic ant species (Tetramorium caespitum); thus this ant may affect native ant richness in urban habitats. Ant composition differed with habitat type, and abundance was lowest in forests. Site characteristics varied with habitat type: forests were larger, had more woody plants, higher woody plant richness, more branches, and leaf litter whereas lots and gardens had more concrete and buildings. Vacant lots had taller herbaceous vegetation, and gardens had higher forb richness, density, and more bare ground. Differences in vegetation did not correlate with ant richness, but several vegetation factors (e.g. patch size, number and size of trees, leaf litter, and amount of concrete and buildings) correlated with differences in ant species composition. Additional factors relating to soil, nests, or microclimatic factors may also be important for urban ant communities. Implications for biodiversity conservation in urban ecosystems are discussed.     

Field and remotely sensed measures of soil and vegetation carbon and nitrogen across an urbanization gradient in the Boston metropolitan area

Understanding the impact of urbanization on terrestrial biogeochemistry is critical for addressing society’s grand challenge of global environmental change. We used field observations and remotely sensed data to quantify the effects of urbanization on vegetation and soils across a 100-km urbanization gradient extending from Boston to Harvard Forest and Worcester, MA. At the field-plot scale, the normalized difference vegetation index (NDVI) was positively correlated with aboveground biomass (AGB) and foliar nitrogen (N) content and negatively correlated with impervious surface fraction. Unlike previous studies, we found no significant relationship between NDVI or impervious surface area (ISA) fraction and foliar N concentration. Patterns in foliar N appeared to be driven more strongly by changes in species composition rather than phenotypic plasticity across the urbanization gradient. For forest and non-residential development, soil nitrogen content increased with urban intensity. In contrast, residential land had consistently high soil N content across the gradient of urbanization. When field observations were scaled-up to the Boston Metropolitan Statistical Area (MSA), we found that soil and vegetation N content were negatively correlated with ISA fraction, an indicator of urban intensity. Our results demonstrated the importance of accounting for the influence of impervious surfaces when scaling field data across urban ecosystems. The combination of field data with remote sensing holds promise for disentangling the complex interactions that drive biogeochemical cycling in urbanizing landscapes. Empirical data that accurately characterize variations in urban biogeochemistry are critical to gain a mechanistic understanding of urban ecosystem function and to guide policy makers and planners in developing ecologically sensitive development strategies.     

Amphibian use of constructed and remnant wetlands in an urban landscape

Urban areas are increasing in number, extent, and human population density worldwide. There is potential to mitigate negative impacts of urbanization to native pond-breeding amphibians by providing habitat in both remnant natural and constructed wetlands. This study examines amphibian use of potential breeding sites in natural and constructed ponds in a large metropolitan area to investigate habitat characteristics that are associated with successful breeding. I surveyed 62 ponds over three breeding seasons in Portland, Oregon, measuring eleven habitat characteristics that may influence their successful breeding: pond depth, nitrate level, aquatic refugia, aquatic vegetation, surrounding vegetation, pond permanence, presence of fish and of introduced bullfrogs, surrounding road density and forest cover, and whether they were constructed or remnant natural ponds. Five of the six native pond-breeding species that occur in the region were regularly found breeding in city ponds. Surrounding forest cover and amount of aquatic vegetation were highly associated with breeding, indicating that preserving and planting vegetation likely benefits urban amphibians. Non-native bullfrogs were not associated with native species richness. Surprisingly, whether a pond was natural or constructed was also only weakly associated with native species breeding, and the trend was towards higher presence for all species in constructed ponds. This indicates that novel, human-dominated areas can provide habitat for these species. Consideration of habitat characteristics associated with breeding success in urban pond management will likely benefit native amphibians in these rapidly expanding landscapes.     

Effects of urban structure on plant species richness in a large European city

Urban areas have developed under anthropogenic impact and hence feature strongly altered environmental conditions that influence today’s plant species distribution. Generally, urban areas are rich in plant species, but on smaller scales richness can vary considerably. Our study aim is to identify areas of high species richness and to assess richness distribution patterns in the city of Hamburg by analyzing a floristic mapping dataset on the scale of 1 km². Differences in plant species richness were analyzed between three urbanization zones. With multiple regression analyses, we tested effects of urban structure, habitat, and environmental conditions on the distribution of species richness measures. Total species richness per 1 km² was 274?±?60 on average and differed only slightly between the urbanization zones. It increased with habitat diversity and decreased with Ellenberg indicator values (EIV) for nutrients (multiple R²?=?0.31). Proportion of non-native species increased with mean annual temperature and decreased with EIVs for moisture (multiple R²?=?0.72), while proportion of endangered species increased with EIVs for moisture and decreased with EIVs for nutrients (multiple R²?=?0.66). Proportion of thermophilic species (multiple R²?=?0.58) could be explained by mean annual temperature. The emerging patterns probably differ from those in other cities due to the central port harboring a particular flora. Besides the expected high proportions of non-native species, high proportions of endangered species were also found in this area. Our results contribute to identifying drivers of biodiversity in cities and can thus be used to develop measures for the conservation of urban biodiversity.     

Fengshui theory in urban landscape planning

The spatial configuration of urban landscapes results from cumulative interactions between human activities and the physical environment. Traditional philosophies and cultural legacies have had important influences on urban development and planning in East Asia. In Seoul, traditional land use practices based on ‘Fengshui’ have significantly contributed to human-mediated patterns of landscape changes, in addition to the role of the socio-economic background (development) and other human activities. The concept of Fengshui was originally founded upon people’s empirical cognition of natural landscape patterns. Recently, however, advanced economic development, westernization and urbanization have been rapidly altering the old traditions of the holistic landscape systems through changing urban planning practices. Since the type, scale, frequency, distribution and spreading pattern of environmental and human disturbances have been changed, a new paradigm for urban landscape planning is necessary to maintain the ecological and cultural integrity of landscapes in Korea. In this paper, we discuss recent concepts and methods of landscape ecology and urban planning from the viewpoint of Fengshui, the traditional land use patterns in Seoul, whose application has so far been restricted only to traditional land evaluation. We conclude that, to maintain the sustainability of the urban landscape, it is necessary to develop a new urban planning framework for the region that is based on the integration between landscape ecology principles with the traditional concepts of Fengshui.     

Butterflies in the city: a review of urban diurnal Lepidoptera

Human migration to urban centers has resulted in diverse environmental disturbances that affect biodiversity. Although urbanization has been highlighted as one of the main drivers of biodiversity endangerment, this topic is still poorly studied in many countries. In order to establish the status quo of the ecology of butterflies in urban centers, we gathered publications focused on urban butterflies (Lepidoptera). We compiled a total of 173 studies from 37 countries and more than 110 urban areas, including published papers and theses (1956–2015). Most papers (69 %) addressed ecological topics, 14 % were focused on biological conservation, and 17 % corresponded to species lists. In summary, most studies revealed a negative impact of urbanization intensity on butterfly diversity (richness and abundance). In fact, we found studies reporting local extinctions due to urbanization, highlighting the causes related to them. The study of charismatic urban wildlife groups, such as butterflies, is a promising field, as there are still important gaps in our comprehension of the ecological patterns and processes that occur in urban areas. Undoubtedly, understanding the response of butterflies to urbanization will aid in the development of urban biodiversity management, planning, and conservation strategies worldwide, which together with knowledge of other wildlife groups and socioeconomic variables, will lead us to more sustainable, livable, and biodiverse cities.     

How small cities affect the biodiversity of ground-dwelling mammals and the relevance of this knowledge in planning urban land expansion in terms of urban wildlife

Fundamental principles regarding urban biodiversity are based on studies conducted in large cities. However we cannot know whether the same phenomena occur in smaller cities or how small cities affect biodiversity. Small cities are an inherent element of urbanization and in the future, most global urban growth is expected to take place in small and medium-sized cities. Understanding the effects of small cities on biodiversity will be an important aspect in planning urban land expansion. Our study examined the effects of a small city on communities of small ground-dwelling mammals on 41 sites arranged in a four step gradient of urbanization. In 6700 trap-days, we caught 2333 individuals comprising 15 species. In the downtown area the same phenomena as those described for large cities were observed: a reduction in species richness and diversity, a decline in the abundance of urban sensitive species and an increase in synurbic species. However, in contrast to large city studies, green areas outside the downtown area did not differ from rural sites in small mammal population parameters. This phenomenon of relatively unchanged fauna outside the downtown area shows that small cities have the potential to maintain a high level of diversity of small ground-dwelling mammals if appropriate planning of further building expansion is implemented. More studies of small cities are needed to better assess their impact on biodiversity. This knowledge can then be applied in better planning for urban wildlife. Generalizations based solely on large city studies are inadequate and may lead to incomplete or inappropriate conservation strategies for small cities.     

Similar effects of residential and non-residential vegetation on bird diversity in suburban neighbourhoods

Estimating the relative importance of vegetation on residential land (gardens, yards, and street-trees) and vegetation on non-residential land (parks and other large green spaces) is important so that competing options for urban conservation planning can be prioritized. We used data from an urban breeding-bird monitoring program to compare the relative effects vegetation on residential land and vegetation on non-residential land (both the amount and type of vegetation at local and landscape scales) on bird species richness and an index of conservation value for the bird community. We then estimated the realised relative benefit of managing the amount of vegetation on these two types of land (i.e., as alternative management options for promoting biodiversity), which might be achieved within the practical limits imposed by human population density. The local effects of increasing residential and non-residential vegetation amount were similar and positive on all measures of bird species richness and conservation value. Non-residential vegetation had an additional landscape-scale influence on bird diversity that residential vegetation did not. Options for managing the amount of non-residential vegetation appear to be more limited by high human population density than for managing the amount of residential vegetation. This suggests that there may be greater realised benefits to bird diversity from managing the amount of vegetation on residential land than from the more common focus of urban planning of managing vegetation on non-residential land.     

Influence of habitat type and distance from source area on bird taxonomic and functional diversity in a Neotropical megacity

The Neotropical region has been subjected to massive urbanization, which poses high risks for some global biodiversity hotspots and losses of ecosystem functions and services. In this study, we investigate how distance from large patches of native forests (source areas) and vegetation (green)/and infrastructure (gray) characteristics affect bird species richness and functional diversity in São Paulo megacity, southeastern Brazil. We analyzed the effects of source areas and green/gray characteristics on species richness and functional diversity (richness, evenness, and divergence) indices. We detected 231 bird species, and our data confirmed our predictions: (1) bird species richness in urbanized habitats was found to be (~?50–85%) lower than in source habitats; (2) species richness and trait composition significantly decreased as the distance from the source area increased, while functional richness was not affected by this metric; and (3) shrub and herbaceous covers and maximum height of trees were positively correlated with species richness and unique functional traits regarding habitat, diet, foraging and nesting strata and dispersal ability of birds in the forest-urban matrix. The number of buildings was negatively correlated with bird species richness and functional richness. Maximum height of buildings caused dramatic declines in functional evenness. Functional divergence was notably lower in sites with high shrub cover. Our study stresses the complexity of vegetation embedded in large Neotropical urban settlements and the need to maintain large protected areas surrounding megacities to mitigate the impacts of urbanization on birds.

     

Bird and mammal use of vernal pools along an urban development gradient

Vernal pools in the northeastern US are of conservation concern primarily because of their role as habitat for specialized pool-breeding amphibians, but their use by birds and mammals may also be of interest, especially from the perspective of the impact of urbanization. We describe camera-trapped wildlife (CTW) at 38 vernal pools along an urban development gradient in greater Bangor, Maine, USA. We detected 20 mammal and 39 bird taxa (29 contacted pool water; 39 detected at >1 site). Land cover type within 1000 m (%), within-pool vegetation (%), and amphibian egg mass numbers explained a substantial portion of the variance (40.8%) in CTW assemblage composition. Submerged vegetation within pools and cover by water and impervious surfaces within 1000 m of pools were key site characteristics defining assemblages. We scored the urban-affiliation of taxa and modeled the relationship between weighted assemblage scores for each site and impervious cover. Impervious cover within 1000 m of pools was positively (p?<?0.01) related to site urban-affiliation scores. Use probability for red fox increased and snowshoe hare decreased with impervious cover at 1000 m. These results indicate that within-pool vegetation and land cover types at 1000 m influenced bird and mammal assemblages that used study pools and greater impervious cover at 100 and 1000 m was correlated with a shift in assemblages from being dominated by urban-avoider to urban-adapted species. We encourage land use planners and managers to consider the influence of land use practices within 1000 m of vernal pools on birds and mammals, especially near amphibian breeding pools.     

Vegetation structure along urban ephemeral streams in southeastern Arizona

Riparian areas in Arizona are being encroached upon by urban developments. This study investigated the impacts of different urban housing densities on riparian vegetation structure along ephemeral streams. Nine sites representing three levels of housing density were selected within the town of Marana, located in southeast Arizona. The housing densities were categorized as high (7–8 houses ha^?1), moderate (2.5–4.5 houses ha^?1), and low (< 1.5 houses ha^?1). Each treatment had three replications. The urban developments were relatively young (less than 15 years). No significant differences were found among the treatments for the tree variables (density, height, mean canopy volume and total canopy volume) or the herbaceous vegetation variables (species richness, percentage of introduced species and percentage of ground cover). However, the shrub variables (mean density, mean height, mean canopy volume, total canopy volume and species richness) showed some significant differences. Shrub density and species richness was significantly greater adjacent to ephemeral channels than just three meters upland. In addition, whitethorn acacia shrubs were significantly taller and larger adjacent to the stream channels in the high and moderate housing density sites than in the low housing density sites. Creosote shrubs showed the opposite trend. Increased runoff in the more heavily urbanized streams may have promoted the growth of the facultative riparian species (whitethorn acacia) but not the non-riparian species (creosote). Overall, in these young developments, vegetation was resilient across the levels of urbanization explored in the study.